Diversions of a NaturalistLankester, E. Ray (Edwin Ray), Sir
History
Diversions of a Naturalist
Lankester, E. Ray (Edwin Ray), Sir
Natural history
The Arthropods differ among themselves in the number of body-rings,
the enlargement or dwindling of certain rings, and the fusion of a
larger or smaller number of the rings to form a composite head, or a
jointless mid-body or hind-body. The successive legs are primarily and
essentially like to one another, and each body-ring, with its pair of
legs, is but a repetition of its fellows. At the same time, in the
different classes included as "Arthropoda" a good deal of difference
has been attained in the structure of the legs, and they have in each
class a different form and character in successive regions of the
body, distinctive of the class, and are sometimes, but not always,
absent from many of the hinder rings. All these Arthropods agree in
having a leg on each side immediately behind the mouth--belonging to a
body-ring, which is fused with others to form the head--very specially
shortened, of great strength and firmness, and shaped so as to be
pulled by a powerful muscle attached to it, against its fellow of the
opposite side, which is similarly pulled. These two stumpy legs form
thus a powerful pair of nippers called "the mandibles." They are jaws,
although they were in the ancestors of the Arthropods merely legs.
These jaw-legs, or leg-jaws, are characteristic of all the crab class,
as well as of the other Arthropods, but no bristle-worm or other animal
has them. The jaws of marine worms are of a totally different nature.
So are the jaws of snails, whelks, and cuttle-fish. Many of the crab
class have not one only, but several, pairs of legs following the
mouth converted into jaws. Thus, if you examine a big shore-crab, or,
better, an edible crab, and a lobster, and a large prawn, you will find
that they all have five pairs of legs converted into short foliaceous
jaws (hence called "foot-jaws"), and overlying the first very strong
pair, or mandibles.
Following these "foot-jaws" you find in a crab or a lobster the great
nipping claws and the four large walking legs--the same in proportion
and shape in crab, lobster, and prawn, much bigger than the foot-jaws.
But the curious thing is that if you set them out and carefully compare
them (for they are not simple jointed limbs, but each has two or even
three diverging stems carried on a basal joint), you will find a
strange and fascinating "likeness in unlikeness," or an agreement of
the parts of which they are built, and yet a difference between all of
them.
The rings of the body to which the jaw-legs and legs are attached are
fused into one unjointed piece. The spine in front of the mouth and
the support of the eyes and the feelers or "antennæ" are fused with
that piece. It forms on the back a great shield--often called "the
head"--which overhangs and is bent down over the sides of this region,
so as to protect the gills, which you can see by cutting away the
overhanging flap.
Public-domain text, read in full here on John Shaqi.
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