Dragons of the Air: An Account of Extinct Flying ReptilesSeeley, H. G. (Harry Govier)
Science
Dragons of the Air: An Account of Extinct Flying Reptiles
Seeley, H. G. (Harry Govier)
Pterosauria
Turning to the Pterodactyles themselves, Von Meyer divided them
naturally into short-tailed and long-tailed. The short-tailed indicated
by the name Pterodactylus he further divided into long-nosed and
short-nosed. The short-nosed genus has since been named Ptenodracon
(Fig. 59, p. 167). The long-tailed group was divided into two types--the
Rhamphorhynchus of the Solenhofen Slate (Fig. 56, p. 161) and the
English form now known as Dimorphodon (Fig. 52, p. 150), which had been
described from the Lias.
The Cretaceous Pterodactyles form a distinct family. So that, believing
the tail to have been short in that group (Fig. 58), there are two
long-tailed as well as two short-tailed families, which were defined
from their typical genera Pterodactylus, Ornithocheirus,
Rhamphorhynchus, and Dimorphodon.
The differences in structure which these animals present are, first: the
big-headed forms from the Lias like Dimorphodon, agree with the
Rhamphorhynchus type from Solenhofen in having a vacuity in the skull
defined by bone, placed between the orbit of the eye and the nostril.
With those characters are correlated the comparatively short bones which
correspond to the back of the hand termed metacarpals, and the tail is
long, and stiffened down its length with ossified tendons. These
characters separate Ornithosaurs with long tails from those with short
tails.
The short-tailed types represented by Pterodactylus and Ornithocheirus
have no distinct antorbital vacuity in the skull defined by bone. The
metacarpal bones of the middle hand are exceptionally elongated, and the
tail, which was flexible in both, appears to have been short. These
differences in the skeleton warrant a primary division of flying
reptiles into two principal groups.
The short-tailed group, which was recognised by De Blainville as
intermediate between Birds and Reptiles, may take the name
Pterodactylia, which he suggested as a convenient, distinctive name. It
may probably be inconvenient to enlarge its significance to comprise not
only the true Pterodactyles originally defined as Pterosauria, but the
newer Ornithostoma and Ornithocheirus which have been grouped as
Ornithocheiroidea.
The second order, in which the wing membrane appears to have had a much
greater extent, in being carried down the hind limbs, where the
outermost digit and metatarsal are modified for its support, has been
named Pterodermata, to include the types which are arranged around
Rhamphorhynchus and Dimorphodon.
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