Dragons of the Air: An Account of Extinct Flying ReptilesSeeley, H. G. (Harry Govier)
Science
Dragons of the Air: An Account of Extinct Flying Reptiles
Seeley, H. G. (Harry Govier)
Pterosauria
In the Cretaceous rocks the scapula presents, in most cases, a different
appearance. The coracoid is an elongated, somewhat triangular bone,
compressed on the outer margin as in birds, but differing alike from
birds and other Pterodactyles in not being prolonged forward beyond the
articulation for the humerus. In these Cretaceous genera, toothed and
toothless alike, the articulation for the upper arm bone truncates the
extremity of the coracoid, so that the bone is less like that of a bird
in this feature. Perhaps it shows a modification towards the crocodilian
direction. The scapula, which unites with the coracoid at about a right
angle, is similarly truncated by the articular surface for the humerus;
but the bone is somewhat expanded immediately beyond the articulation,
and compressed; and instead of being directed backward, it is directed
inward over the ribs to articulate with the neural arches of the early
dorsal vertebræ in the genera found in strata associated with the Chalk.
As the bone approaches this articulation, it thickens and widens a
little, becoming suddenly truncated by an ovate facet, which exactly
corresponds to the transversely ovate impression, concave from front to
back, which is seen in the neural arches of the dorsal vertebræ on which
it fits. This condition is not present in all Cretaceous Pterodactyles.
It does not occur in the Kansas fossil, named by Professor Marsh,
Nyctodactylus. And it appears to be absent from the Pterodactyles of the
English Weald, named Ornithodesmus.
[Illustration: FIG. 39. THE NOTARIUM
An ossification which gives attachment to the scapulæ seen in the
early dorsal vertebra of Ornithocheirus
(From the Cambridge Greensand)]
[Illustration: FIG. 40. RESTORATION OF THE SHOULDER-GIRDLE IN THE
CRETACEOUS ORNITHOCHEIRUS
Showing how the scapulæ articulate with a vertebra and the
articulation of the coracoids with the sternum. The humeral
articulation with the coracoid is unlike the condition shown
in other Ornithosaurs]
There is no approach to this transverse position of the scapulæ among
birds. And while the form of the bones in the older genera of
Ornithosaurs is singularly bird-like, the angular arrangement in this
Cretaceous genus is obtained by closely approximating the articulations
on the sternum, so that the coracoids extend outward as in reptiles,
instead of forward as in birds; and the extremities of the scapulæ
similarly approximate towards each other. This rather recalls the
relative positions of scapula and coracoid among crocodiles. If
crocodile and bird had been primitive types of animals instead of
surviving types, it might almost seem as though there had been a cunning
and harmonious blending of one with the other in evolving this form of
shoulder-girdle.
THE FORE LIMB
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