Dragons of the Air: An Account of Extinct Flying ReptilesSeeley, H. G. (Harry Govier)
Science
Dragons of the Air: An Account of Extinct Flying Reptiles
Seeley, H. G. (Harry Govier)
Pterosauria
what has been termed the lateral carpal as the first metacarpal bone,
which would be as short as that of a bird, but turned in the opposite
direction backward. The first digit would then only carry one phalange,
and would not terminate in a claw, but lie in the line of the tendon
which supports the anterior wing membrane of a bird.
The third bone in the fore-arm of Chrysochloris does not appear to
correspond to a digit. The bone is on the opposite side of the arm to
the similar bone of a Pterodactyle, and therefore cannot be the same
structure in the Golden Mole. The interpretation which makes the pteroid
bone the first digit has the merit of accounting for the fifth digit of
the hand. All the structures of the hand are consistent with this view.
The circumstance that the bone is rarely found in contact with the
radius, but diverging from it, shows that it plays the same part in
stretching the membrane in advance of the arm, that the fifth digit
holds in supporting the larger wing membrane behind the arm.
According to Professor Williston, the American toothless Pterodactyle
Ornithostoma has but a single phalange on the corresponding first toe of
the hind foot, and that bone he describes as long, cylindrical, gently
curved, and bluntly pointed. There is some support for this
interpretation; but I have not seen any English or German Pterodactyles
with only one phalange in the first toe.
The wing in Pterodactyles would thus be stretched between two fingers
which are bent backward, the three intermediate digits terminating in
claws.
THE CARPUS
The wrist bones in the reptilia usually consist of two rows. In
Crocodiles, in the upper row there is a large inner and a small outer
bone, behind which is a lunate bone, the remainder of the carpus being
cartilaginous. Only one carpal is converted into bone in the lower row.
It is placed immediately under the smaller upper carpal. In Chelonians,
the turtle and tortoise group, the characters of the carpus vary with
the family. In the upper row there are usually two short carpals, which
may be blended, under the ulna; while the two under the radius are
commonly united. The lower row is made up of several small bones.
Lizards, too, usually have three bones in the proximal row and five
smaller bones in the distal row.
The correspondence of the distal carpals with the several metacarpal
bones of the middle hand is a well-known feature of the structure of the
wrist.
Von Meyer remarks that the carpus is made up of two rows of small bones
in the Solenhofen Pterodactyles; while in birds there is one row
consisting of two bones. The structure of the carpus is not distinct in
all German specimens; but in the short-tailed Solenhofen genera the
bones in the two rows retain their individuality.
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