Dragons of the Air: An Account of Extinct Flying ReptilesSeeley, H. G. (Harry Govier)
Science
Dragons of the Air: An Account of Extinct Flying Reptiles
Seeley, H. G. (Harry Govier)
Pterosauria
The terminal claw bones are unlike the claws of Birds or Reptiles. They
are compressed from side to side, and extremely deep and strong, with
evidence of powerful attachment for ligaments, so that they rather
resemble in their form and large size the claws of some of the
carnivorous fossil reptiles, often grouped as Dinosauria, such as have
been termed Aristosuchus and Megalosaurus. In the hand of the Ostrich
the first and second digits terminate in claws, while the third is
without a claw. But these claws of the Ostrich and other birds are
slender, curved, and rather feeble organs. In the Archæopteryx, a fossil
bird which agrees with the Pterodactyles in retaining the separate
condition of the metacarpal bones and in having the same number of
phalanges in two of the fingers of the fore limb, the terminal claws are
rather more compressed from side to side, and stronger than in the
Ostrich, but not nearly so strong as in the Pterodactyle. The
Archæopteryx differs from the Pterodactyle in having no trace of a wing
finger. The first metacarpal bone is short, as in all birds; and the
first phalange scarcely lengthens that segment of the first digit of the
Bird's hand to the same length as the other metacarpal bones. It
therefore was not bent backward like the first digit in Pterodactyles.
The wing finger, from which the genius of Cuvier selected the scientific
name--Pterodactyle--for these fossils, yields their most distinctive
character. It is a feature which could only be partly paralleled in the
Bat, by making changes of structure which would remove every support to
the wing but the outermost digit of that animal's hand. In the Bat's
hand the membrane for flight is extended chiefly by four diverging
metacarpal bones. There are only two or three phalanges in each digit in
its four wing fingers. In Pterodactyles the metacarpal bones are, as we
have seen, arranged in close contact, and take no part in stretching the
wing.
THE WING FINGER
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