Dragons of the Air: An Account of Extinct Flying ReptilesSeeley, H. G. (Harry Govier)
Science
Dragons of the Air: An Account of Extinct Flying Reptiles
Seeley, H. G. (Harry Govier)
Pterosauria
The hip-girdle of bones differs chiefly from other types in the way in
which those bones, which have sometimes been likened to the marsupial
bones, are conditioned. They may be a pair of triangular bones which
meet in the middle line, so that there is an outer angle like the arm of
a capital Y. Sometimes these triangular bones are blended into a curved,
bow-shaped arch, which in several specimens appears to extend forward
from near the place of articulation of the femur. This is seen in fossil
skeletons at Heidelberg and Munich. It is possible that this position is
an accident of preservation, and that the prepubic bones are really
attached to the lower border of the pubic bones.
Immense as the length of the tail appears to be, exceeding the skull and
remainder of the vertebral column, it falls far short of the combined
length of the phalanges of the wing finger. The power of flight was
manifestly greater in Rhamphorhynchus than in other members of the
group, and all the modifications of the skeleton tend towards adaptation
of the animals for flying. The most remarkable modification of structure
at the extremity of the tail was made known by Professor Marsh in a
vertical, leaf-like expansion in this genus, which had not previously
been observed (p. 161). The vertebræ go on steadily diminishing in
length in the usual way, and then the ossified structures which bordered
the tail bones and run parallel with the vertebræ in all the
Rhamphorhynchus family, suddenly diverge downward and upward at right
angles to the vertebræ, forming a vertical crest above and a
corresponding keel below; and between these structures, which are
identified with the neural spines and chevron bones of ordinary
vertebræ, the membrane extends, giving the extremity of the tail a
rudder-like feature, which, from knowledge of the construction of the
tail of a child's kite, may well be thought to have had influence in
directing and steadying the animal's movements. There are many minor
features in the shoulder-girdle, which show that the coracoid, for
example, was becoming unlike that bone in the Lias, though it still
continues to have a bony union with the elongated shoulder-blade of the
back.
[Illustration: FIG. 56. RESTORATION OF THE SKELETON OF
_RHAMPHORHYNCHUS PHYLLURUS_
From the Solenhofen Slate, partly based upon the skeleton
with the wing membranes preserved]
[Illustration: FIG. 57. RESTORATION OF THE SKELETON OF _SCAPHOGNATHUS
CRASSIROSTRIS_
Published in the _Illustrated London News_ in 1875. In which a tail is
shown on the evidence of the structure of the head and hand]
[Illustration: FIG. 58. SIX RESTORATIONS
1. Ramphocephalus. Stonesfield Slate. John Phillips, 1871
2. Rhamphorhynchus. O. C. Marsh, 1882
3. Rhamphorhynchus. V. Zittel, 1882
4. Ornithostoma. Williston, 1897
5. Dimorphodon. Buckland, 1836. Tail then unknown
6. Ornithocheirus. H. G. Seeley, 1865]
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