It is a remarkable feature in these animals that the middle bones of the foot, termed instep bones or metatarsals, are usually close together, so that the toes diverge from a narrow breadth, as in _P. longirostris_, _P. Kochi_, and other forms; but there also appear to be splay-footed groups of Pterodactyles like the species which have been named _P. elegans_ and _P. micronyx_, in which the metatarsus widens out so that the bones of the toes do not diverge, and that condition characterises the Ptenodracon (_Pterodactylus brevirostris_), to which genus these species may possibly be referred. Nearly all who have studied these animals regard the singularly short-nosed species _P. brevirostris_ as forming a separate genus. For that genus Sömmerring's descriptive name Ornithocephalus, which he used for Pterodactyles generally, might perhaps have been retained. But the name Ptenodracon, suggested by Mr. Lydekker, has been used for these types. [Illustration: FIG. 62. _CYCNORHAMPHUS SUEVICUS_ Skeleton restored from the bones in Fig. 60] [Illustration: FIG. 63. RESTORATION OF SKELETON CYCNORHAMPHUS FRAASI SHOWING THE LIMBS ON THE RIGHT SIDE _From a specimen in the Museum at Stuttgart_] [Illustration: FIG. 64. CYCNORHAMPHUS FRAASI RESTORATION OF THE FORM OF THE BODY] Some of the largest specimens preserved at Stuttgart and Tübingen have been named _Pterodactylus suevicus_ and _P. Fraasii_. They do not approach the species _P. grandis_ in size, so far as can be judged from the fragmentary remains figured by Von Meyer; for what appears to be the third phalange of the wing finger is 7-1/2 inches long, while in these species it is less than half that length, indicating an enormous development of wing, relatively to the length of the hind limb, which would probably refer the species to another genus. _Pterodactylus suevicus_ differs from the typical Pterodactyles in having a rounded, flattened under surface to the lower jaw, instead of the common condition of a sharp keel in the region of the symphysis. The beak also seems flattened and swan-like, and the teeth are limited to the front of the jaw. There appear to be some indications of small nostrils, which look upward like the nostrils of Rhamphorhynchus, but this may be a deceptive appearance, and the nostrils are large lateral vacuities, which are in the position of antorbital vacuities, so that there would appear to be only two vacuities in the side of the head in these animals. The distinctive character of the skeleton in this genus is found in the extraordinary length of the metacarpus and in the complete ossification of the smaller metacarpal bones throughout their length. The metacarpal bones are much longer than the bones of the fore-arm, and about twice the length of the humerus. The first wing phalange is much longer than the others, which successively and rapidly diminish in length, so that the third is half the length of the first. There are