=272. Movement in response to stimuli.=—If we pinch with the
forceps one of the terminal leaflets, or tap it with a pencil, the two
end leaflets fold above the “vein” of the pinna. This is immediately
followed by the movement of the next pair, and so on as shown in fig.
125, until all the leaflets on this pinna are closed, then the stimulus
travels down the other pinnæ in a similar manner, and soon the pinnæ
approximate each other and the leaf then drops downward as shown in
fig. 126. The normal position of the leaf is shown in fig. 124. If we
jar the plant by striking it or by jarring the pot in which it is grown
all the leaves quickly collapse into the position shown in fig. 126.
If we examine the leaf now we see minute cushions at the base of each
leaflet, at the junction of the pinnæ with the petiole, and a larger
one at the junction of the petiole with the stem. We shall also note
that the movement resides in these cushions.
=273. Transmission of the stimulus.=—The transmission of the
stimulus in this mimosa from one part of the plant has been found to be
along the cells of the bast.
=274. Cause of the movement.=—The movement is caused by a sudden
loss of turgidity on the part of the cells in one portion of the
pulvinus, as the cushion is called. In the case of the large pulvinus
at the base of the petiole this loss of turgidity is in the cells of
the lower surface. There is a sudden change in the condition of the
protoplasm of the cells here so that they lose a large part of their
water. This can be seen if with a sharp knife we cut off the petiole
just above the pulvinus before movement takes place. A drop of liquid
exudes from the cells of the lower side.
=275. Paraheliotropism of the leaves of the sensitive plant.=—If
the mimosa plant is placed in very intense light the leaflets will
turn their edges toward the incidence of the rays of light. This is
also true of other plants in intense light, and is _paraheliotropism_.
Transpiration is thus lessened, and chlorophyll is protected from too
intense light.
[Illustration: Fig. 126_a_. Leaf of Venus fly-trap (Dionæa muscipula),
showing winged petiole and toothed lobes.]
[Illustration: Fig. 127. Leaf of Drosera rotundifolia, some of the
glandular hairs folding inward as a result of a stimulus.]
We thus see that variations in the intensity of light have an important
influence in modifying movements. Variations in temperature also exert
a considerable influence, rapid elevation of temperature causing
certain flowers to open, and falling temperature causing them to close.
=276. Sensitiveness of insectivorous plants.=—The Venus fly-trap
(Dionæa muscipula) and the sundew (drosera) are interesting examples of
sensitive plants, since the leaves close in response to the stimulus
from insects.
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