=305. Fertilization.=—Fertilization is accomplished by the
spermatozoids swimming in at the open end of the oogonium, when one of
them makes its way down into the egg and fuses with the nucleus of the
egg.
[Illustration: Fig. 143. Fertilization of vaucheria. _fn_, female
nucleus; _mn_, male nucleus. The different figures show various stages
in the fusion of the nuclei.]
=306. The twin vaucheria (V. geminata).=—Another species of
vaucheria is the twin vaucheria. This is also a common one, and may be
used for study instead of the sessile vaucheria if the latter cannot
be obtained. The sexual organs are borne at the end of a club-shaped
branch. There are usually two oogonia, and one antheridium between them
which terminates the branch. In a closely related species, instead of
the two oogonia there is a whorl of them with the antheridium in the
center.
=307. Vaucheria compared with spirogyra.=—In vaucheria we have a
plant which is very interesting to compare with spirogyra in several
respects. Growth takes place, not in all parts of the thread, but is
localized at the ends of the thread and its branches. This represents a
distinct advance on such a plant as spirogyra. Again, only specialized
parts of the plant in vaucheria form the sexual organs. These are
short branches. Farther there is a great difference in the size of the
two organs, and especially in the size of the gametes, the supplying
gametes (spermatozoids) being very minute, while the receptive gamete
is large and contains all the nutriment for the fertilized egg. In
spirogyra, on the other hand, there is usually no difference in size
of the gametes, as we have seen, and each contributes equally in the
matter of nutriment for the fertilized egg. Vaucheria, therefore,
represents a distinct advance, not only in the vegetative condition of
the plant, but in the specialization of the sexual organs. Vaucheria,
with other related algæ, belongs to a group known as the _Siphoneæ_, so
called because the plants are tube-like or _siphon_-like.
[Illustration: Fig. 143_a_. Botrydium granulatum. _A_, the whole plant;
_B_, swarm spore; _C_, planogametes; _a_, a single gamete; _b_-_e_, two
gametes in process of fusion; _f_, zygote.]
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