Portion of moss plant of Mnium affine, showing two sporogonia from one
branch. Capsule at left has just shed the cap or operculum; capsule at
right is shedding spores, and the teeth are bristling at the mouth.
Next to the right is a young capsule with calyptra still attached; next
are two spores enlarged.]
=516. The fruiting moss plant.=—In fig. 280 is a moss plant “in
fruit,” as we say. Above the leafy stem a slender stalk bears the
capsule, and in this capsule are borne the spores. The capsule then
belongs to the _sporophyte phase_ of the moss plant, and we should
inquire whether the entire plant as we see it here is the sporophyte,
or whether part of it is gametophyte. If a part of it is gametophyte
and a part sporophyte, then where does the one end and the other begin?
If we strip off the leaves at the end of the leafy stem, and make a
longisection in the middle line, we should find that the stalk which
bears the capsule is simply stuck into the end of the leafy stem, and
is not organically connected with it. This is the dividing line, then,
between the gametophyte and the sporophyte. We shall find that here the
archegonium containing the egg is borne, which is a surer way of
determining the limits of the two phases of the plant.
=517. The male and female moss plants.=—The two plants of mnium
shown in figs. 281, 282 are quite different, as one can easily see,
and yet they belong to the same species. One is a female plant, while
the other is a male plant. The sexual organs then in mnium, as in many
others of the mosses, are borne on separate plants. The archegonia are
borne at the end of the stem, and are protected by somewhat narrower
leaves which closely overlap and are wrapped together. They are similar
to the archegonia of the liverworts.
[Illustration: Fig. 281. Female plant (gametophyte) of a moss (mnium),
showing rhizoids below, and the tuft of leaves above which protect the
archegonia.]
[Illustration: Fig. 282. Male plant (gametophyte) of a moss (mnium)
showing rhizoids below and the antheridia at the center above
surrounded by the rosette of leaves.]
The male plants of mnium are easily selected, since the leaves at the
end of the stem form a broad rosette with the antheridia, and some
sterile threads packed closely together in the center. The ends of the
mass of antheridia can be seen with the naked eye, as shown in fig.
282. When the antheridia are ripe, if we make a section through a
cluster, or if we merely tease out some from the end with a needle in
a drop of water on the slide, then prepare for examination with the
microscope, we can see the form of the antheridia. They are somewhat
clavate or elliptical in outline, as seen in fig. 284. Between them
there stand short threads composed of several cells containing
chlorophyll grains. These are sterile threads (paraphyses).
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