=534.= In the front of the sporangium is another peculiar group
of cells. Two of the longer ones resemble the lips of some creature,
and since the sporangium opens between them they are sometimes termed
the lip cells. These lip cells are connected with the upper end of the
annulus on one side and with the upper end of the stalk on the other
side by thin-walled cells, which may be termed connective cells, since
they hold each lip cell to its part of the opening sporangium. The
cells on the side of the sporangium are also thin-walled. If we now
examine a sporangium from the back, or dorsal edge as we say, it will
appear as in the left-hand figure. Here we can see how very prominent
the annulus is. It projects beyond the surface of the other cells of
the sporangium. The spores are contained inside this case.
=535. Opening of the sporangium and dispersion of the spores.=—If
we take some fresh fruiting leaves of the Christmas fern, or of any one
of many of the species of the true ferns just at the ripening of the
spores, and place a portion of it on a piece of white paper in a dry
room, in a very short time we shall see that the paper is being dusted
with minute brown objects which fly out from the leaf. Now if we take
a portion of the same leaf and place it under the low power of the
microscope, so that the full rounded sporangia can be seen, in a short
time we note that the sporangium opens, the upper half curls backward
as shown in fig. 294, and soon it snaps quickly, to near its former
position, and the spores are at the same time thrown for a considerable
distance. This movement can sometimes be seen with the aid of a good
hand lens.
[Illustration: Fig. 290. Four pinnæ of adiantum, showing recurved
margins which cover the sporangia.]
[Illustration: Fig. 291. Section through sorus of Polypodium vulgare
showing different stages of sporangium, and one multicellular capitate
hair.]
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