=538. The movements of the sporangium can take place in old and dried
material.=—If we have no fresh material to study the sporangium
with, we can use dried material, for the movements of the sporangia can
be well seen in dried material, provided it was collected at about the
time the sporangia are mature, that is at maturity, or soon afterward.
We take some of the dry sporangia (or we may wash the glycerine off
those which we have just studied) and mount them in water, and quickly
examine them with a microscope. We notice that in each cell of the
annulus there is a small sphere of some gas. The water which bathes the
walls of the annulus is absorbed by some substance inside these cells.
This we can see because of the fact that this sphere of gas becomes
smaller and smaller until it is only a mere dot, when it disappears in
a twinkling. The water has been taken in under such pressure that it
has absorbed all the gas, and the farther pressure in most cases closes
the partly opened sporangium more completely.
=539.= Now we should add glycerine again and draw out the water,
watching the sporangia at the same time. We see that the sporangia
which have opened and snapped once will do it again. And so they may
be made to go through this operation several times in succession. We
should now note carefully the annulus, that is after the sporangia have
opened by the use of glycerine. So soon as they have snapped in the
glycerine we can see those minute spheres of gas again, and since there
was no air on the outside of the sporangia, but only glycerine, this
gas must, it is reasoned, have been given up by the water before it was
all drawn out of the cells.
=540. The common polypody.=—We may now take up a few other ferns
for study. Another common fern is the polypody, one or more species of
which have a very wide distribution. The stem of this fern is also not
usually seen, but is covered with the leaves, except in the case of
those species which grow on the surface of rocks. The stem is slender
and prostrate, and is covered with numerous brown scales. The leaves
are pinnate in this fern also, but we find no difference between the
fertile and sterile leaves (except in some rare cases). The fruit dots
occupy much the same positions on the under side of the leaf that they
do in the Christmas fern, but we cannot find any indusium. In the place
of an indusium are club-shaped hairs as shown in fig. 291. The enlarged
ends of these clubs reaching beyond the sporangia give some protection
to them when they are young.
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