=567.= It is quite interesting to note the form of these changed
leaves carefully, to see how this change has affected the pinnæ and the
sporangia. We note that the tip of the leaf as well as the tips of all
the pinnæ are more expanded than the basal portions of the same.
This is due to the fact that the tip of the leaf develops later
than the basal portions. At the time the stimulus to the change in
the development of the fertile leaves reached them they were partly
formed, that is the basal parts of the fertile leaves were more or less
developed and fixed and could not change. Those portions of the leaf,
however, which were not yet completely formed, under this stimulus, or
through correlation of growth, are incited to vegetative growth, and
expand more or less completely into vegetative leaves.
=568. The sporangia decrease as the fertile leaf expands.=—If
we now examine the sporangia on the successive pinnæ of a partly
transformed leaf we find that in case the lower pinnæ are not changed
at all, the sporangia are normal. But as we pass to the pinnæ which
show increasing changes, that is those which are more and more
expanded, we see that the number of sporangia decrease, and many of
them are sterile, that is they bear no spores. Farther up there are
only rudiments of sporangia, until on the more expanded pinnæ sporangia
are no longer formed, but one may still see traces of the indusium.
On some of the changed leaves the only evidences that the leaf began
once to form a fertile leaf are the traces of these indusia. In some of
these cases the transformed leaf was even larger than the sterile leaf.
=569. The ostrich fern.=—Similar changes were also produced in
the case of the ostrich fern, and in fig. 319 is shown at the left a
normal fertile leaf, then one partly changed, and at the right one
completely transformed.
=570. Dimorphism in tropical ferns.=—Very interesting forms
of dimorphism are seen in some of the tropical ferns. One of these
is often seen growing in plant conservatories, and is known as the
staghorn fern (Platycerium alcicorne). This in nature grows attached to
the trunks of quite large trees at considerable elevations on the tree,
sometimes surrounding the tree with a massive growth. One kind of leaf,
which may be either fertile or sterile, is narrow, and branched in a
peculiar manner, so that it resembles somewhat the branching of the
horn of a stag. Below these are other leaves which are different in
form and sterile. These leaves are broad and hug closely around the
roots and bases of the other leaves. Here they serve to catch and
retain moisture, and they also catch leaves and other vegetable matter
which falls from the trees. In this position the leaves decay and then
serve as food for the fern.
[Illustration: Fig. 320. Ostrich fern, showing one normal sporophyll,
one partly transformed, and one completely transformed.]
CHAPTER XXIX.
HORSETAILS.
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