=577. Gametophyte of equisetum.=—The spores of equisetum have
chlorophyll when they are mature, and they are capable of germinating
as soon as mature. The spores are all of the same kind as regards size,
just as we found in the case of the ferns. But they develop prothallia
of different sizes, according to the amount of nutriment which they
obtain. Those which obtain but little nutriment are smaller and develop
only antheridia, while those which obtain more nutriment become larger,
more or less branched, and develop archegonia. This character of an
independent prothallium (gametophyte) with the characteristic sexual
organs, and the also independent sporophyte, with spores, shows the
relationship of the horsetails with the ferns. We thus see that these
characters of the reproductive organs, and the phases and fruiting of
the plant, are more essential in determining relationships of plants
than the mere outward appearances.
CHAPTER XXX.
CLUB MOSSES.
[Illustration: Fig. 326. Lycopodium clavatum, branch bearing two
fruiting spikes; at right sporophyll with open sporangium; single spore
near it.]
=578.= What are called the “club mosses” make up another group
of interesting plants which rank as allies of the ferns. They are not
of course true mosses, but the general habit of some of the smaller
species, and especially the form and size of the leaves, suggest a
resemblance to the larger of the moss plants.
=579. The clavate lycopodium.=—Here is one of the club mosses
(fig. 326) which has a wide distribution and which is well entitled to
hold the name of club because of the form of the upright club-shaped
branches. As will be seen from the illustration, it has a prostrate
stem. This stem runs for considerable distances on the surface of the
ground, often partly buried in the leaves, and sometimes even buried
beneath the soil. The leaves are quite small, are flattened-awl-shaped,
and stand thickly over the stem, arranged in a spiral manner, which
is the usual arrangement of the leaves of the club mosses. Here and
there are upright branches which are forked several times. The end of
one or more of these branches becomes produced into a slender upright
stem which is nearly leafless, the leaves being reduced to mere scales.
The end of this leafless branch then terminates in one or several
cylindrical heads which form the club.
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