=644. Gynœcium.=—Next above the stamens and at the center of the
flower is a stout, angular, ovate body which terminates in three long,
slender, curved points. This is the pistil, and at present the only
suggestion which it gives of belonging to the leaf series is the fact
that the end is divided into three parts, the number of parts in each
successive whorl of members of the flower. If we cut across the body of
this pistil and examine it with a low power we see that there are three
chambers or cavities, and at the junction of each the walls suggest to
us that this body may have been formed by the infolding of the margins
of three leaf-like members, the places of contact having then become
grown together. We see also that from the incurved margins of each
division of the pistil there stand out in the cavity oval bodies. These
are the _ovules_. Now the ovules we have learned from our study of the
gymnosperms are the _sporangia_ (here the macrosporangia). It is now
more evident that this curious body, the pistil, is made up of three
leaf-like members which have fused together, each member being the
equivalent of a sporophyll (here the macrosporophyll). This must be a
fascinating observation, that plants of such widely different groups
and of such different grades of complexity should have members formed
on the same plan and belonging to the same series of members, devoted
to similar functions, and yet carried out with such great modifications
that at first we do not see this common meeting ground which a
comparative study brings out so clearly.
[Illustration: Fig. 381. Abnormal trillium. The nine parts of the
perianth are green, and the outer whorls of stamens are expanded into
petal-like members.]
[Illustration: Fig. 382. Transformed stamen of trillium showing anther
locules on the margin.]
=645. Transformations of the flower of trillium.=—If anything
more were needed to make it clear that the parts of the flower of
trillium belong to the leaf series we could obtain evidence from the
transformations which the flower of trillium sometimes presents. In
fig. 381 is a sketch of a flower of trillium, made from a photograph.
One set of the stamens has expanded into petal-like organs, with the
anther sacs on the margin. In fig. 380 is shown a plant of Trillium
grandiflorum in which the pistil has separated into three distinct and
expanded leaf-like structures, all green except portions of the margin.
Dentaria.
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