=693. (2d) Shoots without foliage leaves.=—These are subterranean
or aerial. Nearly all subterranean shoots have also aerial shoots,
the latter being for the display of foliage leaves (foliage-shoots),
and also for the display of flowers (flower-shoots). The subterranean
kinds bear scale leaves, i.e., the leaves not having a light relation
are reduced in size, being small, and they lack chlorophyll. Examples
are found in Solomon’s seal, mandrake (fig. 413_a_), etc. Here the
scale leaves are on the bud at the end of the underground stem from
which the foliage shoot arises. Aerial shoots which lack foliage
leaves are the dodder, Indian-pipe-plant, beech drops, etc. These
plants are saprophytes or parasites (see Chapter IX). Deriving their
carbohydrate food from other living plants, or from humus, they do not
need green leaves. The leaves have, therefore, probably been reduced
in size to mere scales, and accompanying this there has been a loss of
the chlorophyll. Other interesting examples of aerial shoots without
foliage leaves are the cacti where the stem has assumed the leaf
function and the leaves have become reduced to mere spines. The various
modifications which shoots have undergone accompanying a change in
their leaf relation will be discussed under stems in Chapter XXXIX.
=694. (3d) Floral shoots.=—The floral shoot is the part of the
plant bearing the flower. As interpreted here it may consist of but a
single flower with its stalk, as in Trillium, mandrake, etc., or of
the clusters of flowers on special parts of the stem, termed flower
clusters, as the _catkin_, _raceme_, _spike_, _umbel_, _head_, etc. In
the floral shoot as thus interpreted there are several peculiarities to
observe which distinguish it from the foliage shoot and adapt it to its
life relations.
The floral shoot in many respects is comparable to the foliage shoot,
as seen from the following peculiarities:
(1) It usually possesses, beside the flowers, small
green leaves which are in fact foliage though they are
very much reduced in size, because the function of
the shoot as a foliage shoot is subordinated to the
function of the floral shoot. These small leaves on the
floral shoot are termed _bracts_.
(2) It may be (_a_) unbranched, when it would consist
of a single flower, or (_b_) branched, when there would
be several to many flowers in the flower cluster.
(3) The flower bud has the same origin on the shoot
as the leaf bud; it is either terminal or axillary, or
both.
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