=698. Sclerenchyma, or stone-tissue.=—This is also a
strengthening tissue and consists of cells which do not taper at the
ends and the walls are evenly thickened, sometimes so thick that the
inside (lumen) of the cell has nearly disappeared. Usually such cells
contain no living contents at maturity. Sclerenchyma is very common in
the hard parts of nuts, and underneath the epidermis of stems and
leaves of many plants, as in the underground stems of the bracken fern,
the leaves of pines (fig. 415), etc.
[Illustration: Fig. 414. Transverse section of portion of tomato stem.
_ep_, epidermis; _ch_ chlorophyll-bearing cells; _co_, collenchyma;
_cp_, parenchyma.]
[Illustration: Fig. 415. Margin of leaf of Pinus pinaster, transverse
section, _c_, cuticularized layer of outer wall of epidermis;
_i_, inner non-cuticularized layer; _c´_, thickened outer wall of
marginal cell; _g_, _i´_, hypoderma of elongated sclerenchyma; _p_,
chlorophyll-bearing parenchyma; _pr_, contracted protoplasmic contents.
×800. (After Sachs.)]
[Illustration: Fig. 416. Section through a lenticel of Betula alba
showing stoma at top, phellogen below producing rows of flattened
cells, the cork. (After De Bary.)]
=699. Cork.=—In many cases there is a development of “cork”
tissue underneath the epidermis. Cork tissue is developed by repeated
division of parenchyma cells in such a way that rows of parallel cells
are formed toward the outside. These are in distinct layers, soon lose
their protoplasm and die; there are no intercellular spaces and the
cells are usually of regular shape and fit close to each other. In
some plants the cell walls are thin (cork oak), while in others they
are thickened (beech). The tissue giving rise to cork is called “cork
cambium,” or phellogen, and may occur in other parts of the plant. For
example, where plants are wounded the living exposed parenchyma cells
often change to cork cambium and develop a protective layer of cork.
The walls of cork cells contain a substance termed _suberin_, which
renders them nearly waterproof.
=700. Lenticels.=—These are developed quite abundantly underneath
stomates on the twigs of birch, cherry, beech, elder, etc. The
phellogen underneath the stoma develops a cushion of cork which presses
outward in the form of an elevation at the summit of which is the stoma
(fig. 416). The lenticels can easily be seen.
2. THE FIBROVASCULAR SYSTEM.
=701. Fibrous tissue.=[39]—This consists of thick-walled cells,
usually without living contents which are elongated and taper at the
ends so that the cells, or fibers, overlap. It is common as one of the
elements of the vascular bundles, as wood fibers and bast fibers.
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