=715. Columnar type.=—The columnar type of stem may be simple or
branched. When branching occurs the branches are usually small and in
general subordinate to the main axis. The sunflower (Helianthus annuus)
is an example. The foliage part is mainly simple. The main axis remains
unbranched during the larger part of the growth-period. The principal
flowerhead terminates the stem. Short branches bearing small heads
then arise in the axils of a few of the upper leaves. In dry, poor
soil, or where other conditions are unfavorable, there may be only the
single terminal flowerhead, when the stem is unbranched. The mullein
is another columnar stem. The foliage part is rarely branched, though
branches sometimes occur where the main axis has become injured or
broken. The flower stem is terminal. The corn plant and the Easter lily
are good illustrations also of the columnar stem.
Among trees the Lombardy poplar (Populus fastigiata) is an excellent
example of the columnar type. Though this is profusely branched, the
branches are quite slender and small in contrast with the main axis,
unless by some injury or other cause two large axes may be developed.
As the technical name indicates, the branching is fastigiate, i.e., the
branches are crowded close together and closely surround the central
axis. The royal palm and some of the tree ferns have columnar, simple
stems, but the large, wide-spreading leaves at the top of the stem give
the plant anything but a cylindrical habit. Some cedars and arbor-vitæ
are also columnar.
The advantages of the columnar habit of stem are three: (1) That the
plant stands above other neighboring ones of equal foliage area and
thus is enabled to obtain a more favorable light relation; (2) where
large numbers of plants of the same species are growing close together,
they can maintain practically the same habit as where growing alone;
(3) the advantage gained by other types in their neighborhood in less
shading than if the type were spreading. The cylindrical type can,
therefore, grow between other types with less competition for existence.
[Illustration: Fig. 422. Cylindrical stem of mullein.]
[Illustration: Fig. 423. Conical type of larch.]
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