Next within the bark we find the so-called “woody” portion of the
twig. This is strong and resistant to both breaking and cutting. The
microscope shows it to be composed of the ordinary already known woody
elements,[43] wood fibers, for strengthening purposes, pitted and spiral
vessels as conducting tissue; and intermixed with these some living
parenchyma cells. A cross-section of the stem also shows narrow radial
lines through the wood. These are pith-rays, composed of vertical
plates of living parenchyma cells. These cells, unlike the others in
the wood, are elongated radially, not vertically. The height of the
pith-rays as well as their thickness varies with the species studied.
In the older trunk only the outer portion, a few inches in thickness,
remains light-colored and fresh, and is called sap-wood. The inner
wood is usually darker and harder, and is termed heart-wood. Living
parenchyma cells, in general, are present only in the sap-wood, and
in this almost solely the ascent of sap occurs. Dyestuffs and other
substances are frequently deposited in the walls of the heart-wood.
The third region occupying the center of the twig is the pith. This is
composed ordinarily of angular, little elongated, parenchyma cells,
when mature mostly without cell-contents and filled with air. The pith
region in different trees is quite diversified. It may be hollow,
chambered, contain scattered thick-walled cells, have woody partitions,
or rarely be entirely thick-walled.
The nature of the woody ring is rather perplexing at first; but
its origin is simple. We may conceive that it has developed from a
stem-type like the sunflower, in which the bundles, though separate,
are connected by a continuous cambium ring. In the woody twigs the
numerous bundles are closely packed together, and only separated by
the primary pith-rays extending from the pith to the cortex. Other
secondary pith-rays are produced within each bundle, but they usually
extend only part way from the cortex to the pith. The wood represents
the xylem of the bundle, and the sieve tubes of the bark, the phloem.
=744. Growth in thickness.=—Although the year’s growth does not
increase in length after the first season has passed, it does increase
in diameter very much. From the size of an ordinary little twig it may
at length become a large tree trunk several feet in thickness. Only a
portion of the first year’s growth is produced by the growing point.
All the rest is a product of the cambium, a cylinder of wood being
added to the exterior of the old wood each season. The cambium, here,
as in the sunflower, lies between the phloem and the xylem, forming a
cylinder entirely around the stem. In spring, when active, it becomes
soft and delicate, thus enabling one to easily strip off the bark from
some trees, such as willow, etc., at that season.
Public-domain text, read in full here on John Shaqi.
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