It has been shown by experiment that when sunlight passes through
red colors the temperature is slightly increased, and it has been
suggested that this may be of protection to the living substance which
has ceased working and is in danger of injury from cold. There does
not seem to be much ground for this suggestion, however. It certainly
could not protect the protoplasm of the leaf at night when the cold
is more intense, and during the day would only aggravate matters
by supplying an increased amount of heat, since extremes of heat
and cold in alternation are more harmful to plant life than uniform
cold. Especially would this be the case in alpine climates where the
alternation of heat and cold between day and night is extreme, and
brilliancy of the colors of alpine plants is well known. It seems
more reasonable to suppose that the red color acts as a screen, as
the chlorophyll is disappearing, to protect from the injurious action
of light, certain organic substances which are to be transferred back
from the leaf to the stem for winter storage. So in the case of many
stems in the spring or early summer when the young leaves often have
a reddish color, it is likely that it acts as a screen to protect the
living substance from the strong light at that season of the year until
the chlorophyll screen, which is weak in young leaves, becomes darker
in color and more effective, when the red color often disappears.
=753. Function of foliage leaves.=—In general the function of
the foliage leaf as an organ of the plant is fivefold (see Chapters
IV, VII, VIII, XI), (1) that of carbon dioxide assimilation or
_photosynthesis_, (2) that of transpiration, (3) that of the synthesis
of other organic compounds, (4) that of respiration, and (5) that of
assimilation proper, or the making of new living substance. While none
of these functions are solely carried on in the leaf, it is the chief
seat of the first three of these processes, its form, position, and
structure being especially adapted to the purpose. Assimilation proper,
as well as respiration, probably take place equally in all growing or
active parts.
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