=764.= (2) =Protective covering.=—_Epidermis and
cuticle._—The walls of the epidermal cells are much thickened in some
plants. Sometimes this thickening occurs in the outer wall, or both
walls may be thickened. Variation in this respect as well as the extent
of the thickening occur in different plants and are often correlated
with the extremes of conditions which they serve to meet. The cuticle,
a waxy exudation from the thick wall of the epidermis of many leaves,
also serves as a protection against too great loss of water, or against
the leaf becoming saturated with water during rains. The cabbage,
carnation, etc., have a well-developed cuticle. The effect of the
cuticle in shedding water can be nicely shown by spraying water on a
cabbage leaf or by immersing it in water. Sunken stomata also retard
the loss of water vapor.
_Covers of hair or scales._—In many leaves certain of the cells of the
epidermis grow out into the form of hairs or scales of various forms,
and they serve a variety of purposes. When the hairs form a felt-like
covering as in the common mullein, some antennarias, etc., they lessen
the loss of water vapor because the air-currents close to the surface
of the leaf are retarded. Spines (see the thistles, etc.) also afford a
protection against certain animals.
=765.= (3) =Reduction of surface.=—Reduction of leaf surface
is brought about in a variety of ways. There are two general modes:
(1st) Reduction of surface along with reduction of mass; (2d) Reduction
of surface inversely as the mass. Examples of the first mode are
seen in the dissected leaves of many aquatic plants. In this finely
dissected condition the mass of the leaf substance is much reduced as
well as the leaf surface, but the leaf is less liable to be injured
by movement of the water. In addition it has already been pointed out
that lobed and divided aerial leaves are much less liable to injury
from violent movements of the air, than if a leaf with the same general
outline were entire. The needle leaves of the conifers are also
examples, and they show as well structural provisions for protection
in the thick, hard cell-walls of the epidermis. To offset the reduced
surface there are numerous crowded leaves. Reduction of surface
inversely as the mass, i.e., the mass of the leaf may not be reduced
at all, or it may be more or less increased. In other words, there is
less leaf surface in proportion to the mass of leaf substance. It is
probable in many cases, example: the crowded, overlapping small scale
leaves of the juniper, arbor-vitæ, cypress, cassiope, pyxidanthera,
etc., that there has been a reduction in the size of the leaf, and at
the same time an increase in thickness. This with the crowding together
of the leaves and their thick cell-walls greatly lessens the radiation
of moisture and heat, thus protecting the leaves both in dry and cold
weather. The succulents, like “live-forever,” have a small amount of
surface in proportion to the mass of the leaf.
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