The statement that leaves usually face the light at right angles is to
be taken as a generalized one. The source of the strongest illumination
varies on different days and again at different times of the day.
On cloudy days the zenith is the source of strongest illumination.
The horizontal position of a leaf, where there are no intercepting
lateral or superior objects would receive its strongest light rays
perpendicular to its surface. The fact is, however, that leaves on the
same stem, because of taller or shorter adjacent stems, are so situated
that the rays of greatest illuminating power are directed at some angle
between the zenith and horizon. Many leaves, then, which may have
their upper sides facing the general source of strongest illumination,
do not necessarily face the sun, and they are thus protected from
possible injury from intense light and heat because the direct rays of
sunlight are for the most part oblique. This does not apply, of course,
to those leaves which “follow the sun” during the day. Their specific
constitution is such that intense illumination is beneficial.
The leaf is adjusted as well as may be in different species of varying
constitution, and under different conditions, to a certain balance in
its relation to the factors concerned. The problem then is to interpret
from this point of view the positions and grouping of leaves. Because
of the specific constitution of different plants, and because of a
great variety of conditions in the environment, we see that it is a
more or less complex question.
[Illustration: Fig. 438. Sunflower with young head turned toward morning
sun.]
Public-domain text, read in full here on John Shaqi.
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