The _ovules_ are within the ovary and are arranged in different ways
in different plants. The pollen grain (or better pollen-spore =
microspore), after it has been transferred to the stigma, “germinates,”
and the pollen tube grows down through the tissue of the stigma and
style, or courses down the stylar canal until it reaches the ovule.
Here it usually enters the ovule (macrosporangium) at the _micropyle_
(in some of the ament-bearing plants it enters at the _chalaza_), and
the sperm cells are emptied into the embryo sac in the interior of the
ovule.
=810. Fertilization.=—One of the sperms unites with the egg
in the embryo sac. This is _fertilization_, and from the fertilized
egg the young embryo is formed still within the ovule. _Double
fertilization_,—the other sperm cell sometimes unites with one or both
of the “polar” nuclei which have united to form the “definitive” or
“endosperm” nucleus. As a result of fertilization, the embryo plant is
formed within the ovule, the coats of which enlarge by growth forming
the seed coats, and altogether forming the seed. (See Chapters XXXIV,
XXXV, XXXVI.)
II. Kinds of Flowers.
=811. Absence of certain flower parts.=—The _complete_ flower
contains all the four series of parts. When any one of the series of
parts is lacking, the flower is said to be _incomplete_. Where only one
series of the floral envelopes is present the flowers are said to be
_apetalous_ (the petals are absent), examples: elm, buckwheat, etc.
Flowers which lack both floral envelopes are _naked_. When pistils are
absent but stamens are present the flowers are _staminate_, whether
floral envelopes are present or not; and so when stamens are absent and
pistils present the flower is _pistillate_. If both stamens and pistils
are absent the flower is said to be _sterile_ or _neutral_ (snowball,
marginal or showy flowers in hydrangea). Flowers with both stamens and
pistils, whether or not they have floral envelopes, are _perfect_ (or
hermaphrodite), so if only one of these sets of _essential organs_
of the flower is present the flower is _imperfect_, or _diclinous_.
Sometimes the imperfect, or diclinous, flowers are on the same plant,
and the plant is said to be _monœcious_ (of one household). When
staminate flowers are on certain individual plants, and the pistillate
flowers of the same species are on other individuals, the plant is
_diœcious_ (or of two households). When some of the flowers of a plant
are diclinous and others are perfect, they are said to be _polygamous_.
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