Two different modes of inflorescence are usually recognized in
the arrangement of flowers on the stem. (1) The _corymbose_, or
_indeterminate inflorescence_ (also indefinite inflorescence), in
which the flowers arise from axillary buds, and the terminal bud may
continue to grow. (2) The _cymose_ or _determinate inflorescence_ (also
_definite inflorescence_) in which the flowers arise from terminal
buds. This arrests the growth of the shoot in length.
There are several advantages to the plant in the different modes of
inflorescence, chief among which is the massing of the flowers, thus
increasing the chances for effective pollination.
A. FLOWER CLUSTERS WITH INDETERMINATE INFLORESCENCE.
=818. The simplest mode of indeterminate inflorescence= is where
the flowers arise in the axils of normal foliage leaves, while the
terminal bud, as in the florist’s smilax, the bellwort, moneywort,
apricot, etc., continues to grow. The flowers are _solitary_ and
_axillary_. In other cases which are far more numerous, the flowers are
associated into more or less definite clusters in which are a number
of recognizable types. The word type used in this sense, it should be
understood, does not refer to an original structure which is the
source of others. It merely refers to a mode of inflorescence which we
attempt to recognize, and about which we group those forms which have
a resemblance to one another. There are many forms of flower clusters
which do not conform to any one of our recognized types, and are very
puzzling. The evolution of the flower clusters has been _natural_, and
we cannot make them all conform to an _artificial_ classification.
These _types_ are named merely as a matter of convenience in the
expression of our ideas. The types usually recognized are as follows:
=819. The raceme.=—The flower-shoot is more or less elongated,
and the leaves are reduced to a minute size termed _bracts_, while the
flowers on lateral axes are solitary in the axils of the bracts. The
reduction in the size of the leaves and the somewhat limited growth
of the shoot in length, makes the flowers more prominent, and brings
them into closer relation than if they were formed in the axils of
the leaves on the ordinary foliage shoot. The choke cherry, currant,
pokeweed, sourwood, etc., are examples of a raceme (fig. 569). In most
plants with the raceme type, while the inflorescence is indeterminate,
and the uppermost flowers (those toward the end of the main shoot)
are younger, still the period of flowering is somewhat restricted
and the raceme stops growing. In a few plants, however, as in the
common “shepherd’s-purse,” the raceme continues to grow throughout the
summer, so that the lower flowers may have ripened their seed while
the terminal portion of the raceme is still growing and producing
new flowers. Compound racemes are formed when by branching of the
flower-shoot there are several racemes in a cluster, as in the false
Solomon’s seal (Smilacina racemosa).
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