=843.= But a modification of the microspore, the pollen tube,
enables the sperm cell to reach the egg-cell. The tube grows through
the nucellus, or first through the tissues of the ovary, deriving
nutriment therefrom.
=844.= But here an important consideration should not escape us.
The pollen grains (microspores) must in nearly all cases first reach
the pistil, in order that in the growth of this tube a channel may be
formed through which the generative cell can make its way to the egg
cell. The pollen passes from the anther locule, then, to the stigma of
the ovary. This process is termed _pollination_.
Pollination.
=845. Self pollination, or close pollination.=—Perhaps very few
of the admirers of the pretty blue violet have ever noticed that there
are other flowers than those which appeal to us through the beautiful
colors of the petals. How many have observed that the brightly colored
flowers of the blue violet rarely “set fruit”? Underneath the soil
or débris at the foot of the plant are smaller flowers on shorter,
curved stalks, which do not open. When the anthers dehisce, they are
lying close upon the stigma of the ovary, and the pollen is deposited
directly upon the stigma of the same flower. This method of pollination
is _self pollination_, or _close pollination_. These small, closed
flowers of the violet have been termed “_cleistogamous_,” because they
are pollinated while the flower is closed, and fertilization takes
place as a result.
But self pollination takes place in the case of some open flowers.
In some cases it takes place by chance, and in other cases by such
movements of the stamens, or of the flower at the time of the
dehiscence of the pollen, that it is quite certainly deposited upon the
stigma of the same flower.
=846. Wind pollination.=—The pine is an example of
wind-pollinated flowers. Since the pollen floats in the air or
is carried by the “wind,” such flowers are _anemophilous_. Other
anemophilous flowers are found in other conifers, in grasses, sedges,
many of the ament-bearing trees, and other dicotyledons. Such plants
produce an abundance of pollen and always in the form of “dust,” so
that the particles readily separate and are borne on the wind.
=847. Pollination by insects.=—A large number of the plants which
we have noted as being anemophilous are monœcious or diœcious, i.e. the
stamens and pistils are borne in separate flowers. The two kinds of
flowers thus formed, the male and the female, are borne either on the
same individual (monœcious) or on different individuals (diœcious). In
such cases cross pollination, i.e. the pollination of the pistil of
one flower by pollen from another, is sure to take place, if it is
pollinated at all. Even in monœcious plants cross pollination often
takes place between flowers of different individuals, so that more
widely different stocks are united in the fertilized egg, and the
strain is kept more vigorous than if very close or identical strains
were united.
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