=901. Physical factors.=—Some of these factors are water, light,
heat, wind, chemical or physical condition of the soil, etc. _Water_
is a very important factor for all plants. Even those growing on land
contain a large percentage of water, which we have seen is rapidly lost
by transpiration, and unless water is available for root absorption
the plant soon suffers, and aquatic plants are injured very quickly by
drying when taken from the water. Excess of soil water is injurious to
some plants. _Light_ is important in photosynthesis, in determining
direction of growth as well as in determining the formation of suitable
leaves in most plants, and has an influence in the structure of the
leaf according as the light may be strong, weak, etc. _Heat_ has great
influence on plant growth and on the distribution of plants. The
growth period for most vegetation begins at 6° C. (= 43° F.), or in
the tropics at 10°-12° C., but a much higher temperature is usually
necessary for reproduction. Some arctic algæ, however, fruit at 1.8° C.
The upper limit favorable for plants in general is 45°-50° C., while
the optimum temperature is below this. Very high temperatures are
injurious, and fatal to most plants, but some algæ grow in hot springs
where the temperature reaches 80°-90° C. Some desert plants are able to
endure a temperature of 70° C., while some flowering plants of other
regions are killed at 45° C. Some plants are specifically susceptible
to cold, but most plants which are injured by freezing suffer because
the freezing is a drying process of the protoplasm (see p. 374). _Wind_
may serve useful purposes in pollination and in aeration, but severe
winds injure plants by causing too rapid transpiration, by felling
trees, by breaking plant parts, by deforming trees and shrubs, and by
mechanical injuries from “sand-blast.” _Ground covers_ protect plants
in several ways. Snow during the winter checks radiation of heat from
the ground so that it does not freeze to so great a depth, and this is
very important for many trees and shrubs. It also prevents alternate
freezing and thawing of the ground, which “heaves” some plants from
the soil. Leaves and other plant remains mulch the soil and check
evaporation of water. The influence of the _chemical condition_ of the
soil is very marked in alkaline areas where the concentration of salt
in the soil permits a very limited range of species. So the physical
and mechanical conditions of the soil influence plants because the
moisture content of the ground is so closely dependent on its physical
condition. Rocky and gravelly soil, other things being equal, is dry.
Clay is more retentive of moisture than sand, and moisture also varies
according to the per cent of humus mixed with it, the humus increasing
the percentage of moisture retained.
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