=204. Structure of the grain of corn.=—Take grains of corn that
have been soaked in water for 24 hours and note the form and difference
in the two sides (in all of these studies the form and structure of
the seed, as well as the stages in germination, should be illustrated
by the student). Make a longisection of a grain of corn through the
middle line, if necessary making several in order to obtain one which
shows the structures well near the smaller end of the grain. Note
the following structures: 1st, the hard outer “wall” (formed of the
consolidated wall of the ovary with the integuments of the ovules—see
Chapters 35 and 36); 2d, the greater mass of starch and other plant
food (the endosperm) in the centre; 3d, a somewhat crescent-shaped body
(the _scutellum_) lying next the endosperm and near the smaller end of
the grain; 4th, the remaining portion of the young embryo lying between
the scutellum and the seed coat in the depression. When good sections
are made one can make out the radicle at the smaller end of the seed,
and a few successive leaves (the plumule) which lie at the opposite
end of the embryo shown by sharply curved parallel lines. Observe the
attachment of the scutellum to the caulicle at the point of junction
of the plumule and the radicle. The scutellum is a part of the embryo
and represents a cotyledon. The endosperm is also called _albumen_, and
such a seed is _albuminous_.
Dissect out an embryo from another seed, and compare with that seen in
the section.
=205.= In the germination of the grain of corn the endosperm
supplies the food for the growth of the embryo until the roots are
well established in the soil and the leaves have become expanded and
green, in which stage the plant has become able to obtain its food from
the soil and air and live independently. The starch in the endosperm
cannot of course be used for food by the embryo in the form of starch.
It is first converted into a soluble form and then absorbed through
the surface of the scutellum or cotyledon and carried to all parts of
the embryo. An enzyme developed by the embryo acts upon the starch,
converting it into a form of sugar which is in solution and can thus
be absorbed. This enzyme is one of the so-called diastatic “ferments”
which are formed during the germination of all seeds which contain food
stored in the form of starch. In some seedlings, this diastase formed
is developed in much greater abundance than in others, for example,
in barley. Examine grains of corn still attached to seedlings several
weeks old and note that a large part of their content has been used up.
The action of diastase on starch is described in Chapter 8.
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