1076. The effort of the three vegetable processes, to separate their
organs from, and perfect them independently of, each other, is in
incessant operation, both from without inwards, as well as from above
downwards, because in both cases is light there, darkness here, dryness
there, humidity here. The cellular tissue, that has finally become
independent in the transverse direction or from without inwards, is
called _bark_, the self-substantial tubular tissue is called _liber_,
the non-dependent tracheal tissue, _wood_.
1. TRACHEAL-SYSTEM, WOOD.
1077. With increased influence of light the tracheal fasciculi also
increase, and form a circle of columns in the parenchyma around the
centre of the plant. Between the column, externally and internally to
the same, is the parenchyma. The more, however, the columns accumulate,
by so much the more does it diminish, and whereas the columns
previously stood singly in the parenchyma, the appearance is now as if
narrow _plates_ only of parenchyma traversed between the columns from
without inwards. Finally the columns predominate to such an extent and
approximate so closely, that the _plates_ almost disappear. They are
now called insertions of cellular tissue, or _medullary rays_. As the
tracheæ convey air, and have thus been more exposed to the process of
oxydation, they generally harden sooner than in other parts.
1078. Around the fasciculi of spiral vessels the cellular tissue also
strives to elongate, and begins at the same time to harden. Such
extended cells, in which the light has almost disappeared, are called
fibres. Indurated tracheæ and fibres are called _Wood_. The wood is
always in the vicinity of the spiral vessels. It is a production
synchronous with the latter.
1079. Only, where spiral vessels are, can genuine wood originate; but
it is not everywhere, where they are found, that woody fibres must be
also present, although the cells extend around all bundles of spiral
vessels. If the degree of oxydation of the cells be slight, they do not
harden, but continue herbaceous in texture. The parenchyma has now been
separated by a circle of fibrous columns into an external and internal,
or peripheric and central. The central parenchyma becomes void of sap
and spongy, because the plant imbibing its nutriment on the surface,
and the air and the light operating thereupon, the processes conduct it
thither. This withered parenchyma is called _pith_, which in accordance
with its origin merits no physiological consideration, nor is worthy
and susceptible of any philosophical construction.
2. TUBULAR SYSTEM, LIBER.
1080. As the plant draws in its nourishment from without, so is the
main proportion of the sap necessarily present in the periphery of the
spiral vessels. The elongated cells in the neighbourhood of the spiral
vessels, and which principally contain sap, are called _Liber_.
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