1280. Every leaf that has been closed in a vesicular or tubular form
is a _follicle_ or _carpel_. There are therefore uni-, bi-, and
tri-locular ovaria, &c. The loculi or cells of the ovarium are none
other than closed carpels. As many therefore as there are of the
former, so many are there of the latter, and vice versâ. The septa or
partition-walls are none other than the involuted edges of the closed
and confluent carpels.
1281. Uni-locular ovaria consist therefore only of one leaf. The legume
is only a compressed carpel.
1282. Every carpel or every cell has its raphé or suture directed
inwards or along the axis of the flower. For the leaves are always so
conjoined that the two halves of the upper or inner side stand counter
to each other.
1283. All other sutures are adventitious and, by their mode of
dehiscence, determine those parts of the ovarium which are called
valves. These sutures are either _dorsal_ or on the back of the
carpel, e. g. _capsula loculicida_; _conjunctive_, where two carpels
abut against each other, e. g. _capsula septicida_; or finally, between
both by the side of the dorsal suture, so that the valve springing up
resembles a shutter, as in many siliquæ or pods.
1284. The columella of the ovary is none other than the internal edge
of the carpels from which the leaf-wall has been freed.
1285. Each carpel-leaf is to be regarded as the common petiole of a
pinnate leaf, upon whose lateral petioles the seeds depend. The seeds
always hang therefore upon the inner angle of the cells.
1286. As the parts of the corolla alternate with the calyx, so do the
carpels or cells of the ovary with the corolla; they stand therefore
opposite the parts of the calyx or are situated in front of them.
1287. The parts of the ovary follow also the uneven series of numbers,
one, three, five. The number two is usually found in irregular, e. g.
the labiate, corollæ.
1288. If few cells be present as parts of the flower, the carpels are
then to be regarded as arrested. In the personate corollæ three are
arrested, but in the papilionaceous, four. The legumen is only a fifth
part of the ovarium.
1289. The development of the carpels stands usually in an inverse
relation to the size of the parts of the corolla. Thus the legume is
situated between the two insignificant petals of the keel, opposite
to the large vexillum; in the Personatæ a carpel is situated in the
fissure of the upper lip; upon the lower lip, consisting of three
lobes, only one carpel is situated, which consequently supplies the
place of four, and is therefore also larger.
1290. The stages of leaf-formation are also displayed in the matured
state of the ovarium. The _radical_ or _scale-leaf_ is repeated in the
_cariopsis_; as in the grasses, orachs, nettles, and such like plants.
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