2204. Where the branchiæ have assumed the laminated form, they are
surrounded by a similarly formed covering or operculum. The shells
of the Mollusca are, as regards their signification, none other than
branchial opercula, or gill-covers; the same holds good also of the
gasteropodal and crustacean shells, and at bottom of every calcareous
and horny covering of the body.
2205. To the same category belong also the opercula of Fishes, and even
their scales. Fundamentally, the whole epidermis is only a product of
respiration or oxydation.
2206. The first saccular or cæcal inversion of the integument, as in
the Leeches, Molluscs, and Snails, is at once a predominance of the
tegumentary, obtained through the medium of the branchial formation,
whereby the skin commences to become a self-substantial organ of
respiration.
2207. In the Scorpions the branchiæ are introverted sacs or cysts, into
which, however, instead of water, air already enters.
2208. This insaccation is converted in the Spiders into more distinct
air-cysts, which finally ramify in the higher insects and become true
tracheæ.
2209. Lastly, the respiratory system obtains the upper hand to such
an extent, that, together with internal air-tubes, external branchial
laminæ are also developed, as in the Molluscs; but in them the tracheæ
obtain the preponderance over the blood-vessels, so that these laminæ
dry up and become wings or fins.
2210. The wings of Insects are branchial laminæ, converted into
air-organs.
2211. The wing-coverings or elytra are branchial opercula, and
correspond to the shells of bivalve Mollusca.
2212. Every insect therefore must properly possess four wings and two
wing-coverings, of which last, however, rudiments only appear to be
left in the nocturnal Lepidoptera.
2213. In the higher organized animals those gills only which are nearer
the head are persistent, the posterior or lateral branchiæ being
gradually arrested.
2214. These lateral branchiæ remain in Fishes as lateral mucous
openings, which constitute the lateral line.
2215. The cervical branchiæ are limited to the number five, which
has already begun to be established in the Crustacea; namely, at the
origins or roots of the five anterior pairs of feet.
2216. The number five probably derives its origin from the vegetable
kingdom, and that indeed from the genesis of the pinnate leaves, so
that one kind of numerical law appears to prevail with respect to this
organ in both organic kingdoms. It is probably also an imitation of
the five organs of sense.
2217. The vessels of the branchiæ of Fishes are accompanied by bony
rings, which correspond to the feet of Crustacea.
2218. All Fishes have, with few exceptions, five branchial arches.
2219. When in Fishes the sarcous system begins to produce viscera, then
the five branchial foramina pass inwards, and only a single respiratory
aperture is left for them in the fleshy body; namely, the external
branchial foramen.
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