Elements of Structural and Systematic Botany: For High Schools and Elementary College CoursesCampbell, Douglas Houghton
Science
Elements of Structural and Systematic Botany: For High Schools and Elementary College Courses
Campbell, Douglas Houghton
Botany -- Textbooks
The growing tissue (cambium), separating the phloem from the wood,
is made up of cells quite like those of the phloem, into which they
insensibly merge, except that their walls are much thinner, as is
always the case with rapidly growing cells. These cells (_B_,
_cam._) are arranged in radial rows and divide, mainly by walls, at
right angles to the radii of the stem. If we examine the inner side
of the ring, the change the cells undergo is more marked. They
become of nearly equal diameter in all directions, and the walls
become woody, showing at the same time distinct stratification (_B_,
_x_).
On examining the xylem, where two growth rings are in contact, the
reason of the sharply marked line seen when the stem is examined
with the naked eye is obvious. On the inner side of this line (_I_),
the wood cells are comparatively small and much flattened, while the
walls are quite as heavy as those of the much larger cells (_II_)
lying on the outer side of the line. The small cells show the point
where growth ceased at the end of the season, the cells becoming
smaller as growth was feebler. The following year when growth
commenced again, the first wood cells formed by the cambium were
much larger, as growth is most vigorous at this time, and the wood
formed of these larger cells is softer and lighter colored than that
formed of the smaller cells of the autumn growth.
The wood is mainly composed of tracheids, there being no vessels
formed except the first year. These tracheids are characterized by
the presence of peculiar pits upon their walls, best seen when thin
longitudinal sections are made in a radial direction. These pits
(Fig. 76, _D_, _p_) appear in this view as double circles, but if
cut across, as often happens in a cross-section of the stem, or in a
longitudinal section at right angles to the radius (tangential),
they are seen to be in shape something like an inverted saucer with
a hole through the bottom. They are formed in pairs, one on each
side of the wall of adjacent tracheids, and are separated by a very
delicate membrane (_F_, _p_, _G_, _y_). These "bordered" pits are
very characteristic of the wood of all conifers.
The structure of the root is best studied in the seedling plant, or
in a rootlet of an older one. The general plan of the root is much
like that of the pteridophytes. The fibro-vascular bundle (Fig. 75,
_M_, _fb._) is of the so-called radial type, there being three xylem
masses (_x_) alternating with as many phloem masses (_ph._) in the
root of the seedling. This regularity becomes destroyed as the root
grows older by the formation of a cambium ring, something like that
in the stem.
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