In this genus it develops
no perivisceral portion, as in other groups, but gives rise solely to
the nephridia and to the reproductive organs. It is probable, though not
certainly proved, that the history of the coelom in other Arthropods is
essentially similar to that of _Peripatus_, allowance being made for the
fact that the nephridial portion does not attain full development in
those forms which are without nephridia in the adult.
With regard to the development of the vascular system, little can be
said here, except that it appears to arise from the spaces of the
mesoblastic reticulum. When this reticulum is sparse or so delicate as
to give way in manipulation, these spaces appear to be represented by a
continuous space which in the earliest stages of development is
frequently spoken of as the blastocoel or segmentation cavity. They
acquire special epithelial walls, and form the main trunks and network
of smaller vessels found in animals with a canalicular vascular system,
or the large sinus-like spaces characteristic of animals with a
haemocoelic body-cavity.
Transient embryonic organs.
The existence of a phase at the beginning of life during which a young
animal acquires its equipment by a process of growth of the germ is of
course intelligible enough; such a phase is seen in the formation of
buds, and in the sexual reproduction of both animals and plants. The
remarkable point is that while in most cases this embryonic growth is a
direct and simple process--e.g. animal and plant buds, embryonic
development of plant seeds--in many cases of sexual reproduction of
animals it is not direct, and the embryonic phase shows stages of
structure which seem to possess a meaning other than that of being
merely phases of growth. The fact that these stages of structure through
which the embryo passes sometimes present for a short time features
which are permanent in other members of the same group, adds very
largely to the interest of the phenomenon and necessitates its careful
examination. This may be divided into two heads: (1) in relation to
embryos, (2) in relation to larvae. So far as embryos are concerned, we
shall limit ourselves mainly to a consideration of the Vertebrata,
because in them are found most instances of that remarkable phenomenon,
the temporary assumption by certain organs of the embryo of stages of
structure which are permanent in other members of the same group. As is
well known, the embryos of the higher Vertebrata possess in the
structure of the pharynx and of the heart and vascular system certain
features--namely, paired pharyngeal apertures, a simple tubular heart,
and a single ventral aorta giving off right and left a number of
branches which pass between the pharyngeal apertures--which permanently
characterize those organs in fishes. The skeleton, largely bony in the
adult, passes through a stage in which it is entirely without bone, and
consists mainly of cartilage--the form which it permanently possesses in
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