instances, more or less plausible, of a like retention of ancestral
features by larvae might be mentioned, and it must be conceded that
there are strong reasons for supposing that larvae often retain traces,
more or less complete, of ancestral stages of structure. But this
admission does not carry with it any obligation to accept the widely
prevalent view that larval history can in any way be regarded as a
recapitulation of ancestral history. Far from it, for larvae in
retaining some ancestral features are in no way different from adults;
they only differ from adults in the features which they have retained.
Both larvae and adults retain ancestral features, and both have been
modified by an adaptation to their respective conditions of life which
has ever been becoming more perfect.
The conclusion, then, has been reached, that whereas larvae frequently
retain traces of ancestral stages of adult structure, embryos will
rarely do so; and we are confronted again with the question, How are we
to account for the presence in the embryo of numerous functionless
organs which cannot be explained otherwise than as having been inherited
from a previous condition in which they were functional? The answer is
that the only organs of this kind which have been retained are organs
which have been retained by the larvae of the ancestors after they have
been lost by the adult, and have become in this way impressed upon the
development. As an illustration taken from current natural history of
the manner in which larval characters are in actual process of becoming
embryonic may be mentioned the case of the viviparous salamander
(_Salamander atra_), in which the gills, &c., are all developed but
never used, the animal being born without them. In other and closely
allied species of salamander there is a considerable period of larval
life in which the gills and gill-slits are functional, but in this
species the larval stage, for the existence of which there was a
distinct reason, viz. the entirely aquatic habits of life in the young
state, has become at one stroke embryonic by its simple absorption into
the embryonic period. The view, then, that embryonic development is
essentially a recapitulation of ancestral history must be given up; it
contains only a few references to ancestral history, namely, those which
have been preserved probably in a much modified form by previous larvae.
Law of v. Baer.
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