After the Conquest no Englishman retained estates in Essex of any importance, and the chief lay barons at the time of the Survey were Geoffrey de Mandeville and Aubrey de Vere. The de Veres, earls of Oxford, were continuously connected with the county until the extinction of the title two centuries ago. Pleshey was the stronghold of the Mandevilles, and, although the house became extinct in 1189, its descendants in the female line retained the title of earls of Essex. The Honour of Hatfield Peverel held by Ranulf Peverel after the Conquest escheated to the crown in the reign of Henry I., and in the same reign the fief of Robert Gernon passed to the house of Mountfichet. Essex has always been mainly an agricultural county, and the ordinary agricultural pursuits were carried on at the time of the Domesday Survey, which also mentions salt-making, wine-making, bee-culture and cheese-making, while the oyster fisheries have been famous from the earliest historic times. The woollen industry dates back to Saxon times, and for many centuries ranked as the most important industry. Cloth-weaving was introduced in the 14th century, and in the 16th century Colchester was noted for its "bays and says." Colchester also possessed a valuable leather industry in the 16th century, at which period Essex was considered an exceptionally wealthy and prosperous county; Norden, writing in 1594, describes it as "moste fatt, frutefull, and full of all profitable things." The decline of the cloth industry in the 17th century caused great distress, but a number of smaller industries began to take its place. Saffron-culture and silk-weaving were extensively carried on in the 17th century, and the 18th century saw the introduction of the straw-plait industry, potash-making, calico-printing, malting and brewing, and the manufacture of Roman cement. The county returned four members to parliament in 1290. From 1295 it returned two members for the county and two for Colchester. Maldon acquired representation in 1331 and Harwich in 1604. Under the Reform Act of 1832 the county returned four members in four divisions. Under the Representation of the People Act of 1868 Maldon and Harwich each lost one member, and the county returned six members in three divisions.