It would thus appear that the portion of the sea-floor best suited for
receiving and preserving the most varied assemblage of marine organic
remains is the area in front of the land, to which rivers and currents
bring continual supplies of sediment. The most favourable conditions
for the accumulation of a thick mass of marine fossiliferous strata
will arise when the area of deposit is undergoing a gradual
subsidence. If the rate of depression and that of deposit were equal,
or nearly so, the movement might proceed for a vast period without
producing any great apparent change in marine geography, and even
without seriously affecting the distribution of life over the
sea-floor within the area of subsidence. Hundreds or thousands of feet
of sedimentary strata might in this way be heaped up round the
continents, containing a fragmentary series of organic remains
belonging to those forms of comparatively shallow-water life which had
hard parts capable of preservation. There can be little doubt that
such has, in fact, been the history of the main mass of stratified
formations in the earth's crust. By far the largest proportion of
these piles of marine strata has unquestionably been laid down in
water of no great depth within the area of deposit of terrestrial
sediment. The enormous thickness to which they attain seems only
explicable by prolonged and repeated movements of subsidence,
interrupted, however, as we know, by other movements of a contrary
kind.
Since the conditions for the preservation of organic remains exist
more favourably under the sea than on land, marine organisms must be
far more abundantly conserved than those of the land. This is true
to-day, and has, as far as known, been true in all past geological
time. Hence for the purposes of the geologist the fossil remains of
marine forms of life far surpass all others in value. Among them there
will necessarily be a gradation of importance, regulated chiefly by
their relative abundance. Now, of all the marine tribes which live
within the juxta-terrestrial belt of sedimentation, unquestionably the
Mollusca stand in the place of pre-eminence as regards their aptitude
for becoming fossils. They almost all possess a hard, durable shell,
capable of resisting considerable abrasion and readily passing into a
mineralized condition. They are extremely abundant both as to
individuals and genera. They occur on the shore within tide mark, and
range thence down into the abysses. Moreover, they appear to have
possessed these qualifications from early geological times. In the
marine Mollusca, therefore, we have a common ground of comparison
between the stratified formations of different periods. They have been
styled the alphabet of palaeontological inquiry.
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