Fossil evidence may be made to prove the existence of gaps which are not otherwise apparent. As has been already remarked, changes in organic forms must, on the whole, have been extremely slow in the geological past. The whole species of a sea-floor could not pass entirely away, and be replaced by other forms, without the lapse of long periods of time. If then among the conformable stratified formations of former ages we encounter sudden and abrupt changes in the _facies_ of the fossils, we may be certain that these must mark omissions in the record, which we may hope to fill in from a more perfect series elsewhere. The complete biological contrasts between the fossil contents of unconformable strata are sufficiently explicable. It is not so easy to give a satisfactory account of those which occur where the beds are strictly conformable, and where no evidence can be observed of any considerable change of physical conditions at the time of deposit. A group of strata having the same general lithological characters throughout may be marked by a great discrepance between the fossils above and below a certain line. A few species may pass from the one into the other, or perhaps every species may be different. In cases of this kind, when proved to be not merely local but persistent over wide areas, we must admit, notwithstanding the apparently undisturbed and continuous character of the original deposition of the strata, that the abrupt transition from the one _facies_ of fossils to the other represents a long interval of time which has not been recorded by the deposit of strata. A.C. Ramsay, who called attention to these gaps, termed them "breaks in the succession of organic remains." He showed that they occur abundantly among the Palaeozoic and Secondary rocks of England. It is obvious, of course, that such breaks, even though traceable over wide regions, were not general over the whole globe. There have never been any universal interruptions in the continuity of the chain of being, so far as geological evidence can show. But the physical changes which caused the breaks may have been general over a zoological district or minor region. They no doubt often caused the complete extinction of genera and species which had a small geographical range.