In all speculations of this nature, however, it is necessary to reason
from as wide a basis of observation as possible, seeing that so much of
the evidence is negative. Especially needful is it to bear in mind that
the cessation of one or more species at a certain line among the rocks
of a particular district may mean nothing more than that, onward from
the time marked by that line, these species, owing to some change in the
conditions of life, were compelled to migrate or became locally extinct
or, from some alteration in the conditions of fossilization, were no
longer imbedded and preserved as fossils. They may have continued to
flourish abundantly in neighbouring districts for a long period
afterward. Many examples of this obvious truth might be cited. Thus in a
great succession of mingled marine, brackish-water and terrestrial
strata, like that of the Carboniferous Limestone series of Scotland,
corals, crinoids and brachiopods abound in the limestones and
accompanying shales, but disappear as the sandstones, ironstones, clays,
coals and bituminous shales supervene. An observer meeting for the first
time with an instance of this disappearance, and remembering what he had
read about breaks in succession, might be tempted to speculate about the
extinction of these organisms, and their replacement by other and later
forms of life, such as the ferns, lycopods, estuarine or fresh-water
shells, ganoid fishes and other fossils so abundant in the overlying
strata. But further research would show him that high above the
plant-bearing sandstones and coals other limestones and shales might be
observed, once more charged with the same marine fossils as before, and
still farther overlying groups of sandstones, coals and carbonaceous
beds followed by yet higher marine limestones. He would thus learn that
the same organisms, after being locally exterminated, returned again and
again to the same area. After such a lesson he would probably pause
before too confidently asserting that the highest bed in which we can
detect certain fossils marks their final appearance in the history of
life. Some breaks in the succession may thus be extremely local, one set
of organisms having been driven to a different part of the same region,
while another set occupied their place until the first was enabled to
return.
Public-domain text, read in full here on John Shaqi.
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