The thorax is composed of three segments; each bears a pair of jointed
legs, and in the vast majority of insects the two hindmost bear each a
pair of wings. From these three pairs of thoracic legs comes the
name--Hexapoda--which distinguishes the class. And the wings, though not
always present, are highly characteristic of the Hexapoda, since no
other group of the Arthropoda has acquired the power of flight. In the
more generalized insects the abdomen evidently consists of ten segments,
the hindmost of which often carries a pair of tail-feelers, (cerci or
cercopods) and a terminal anal segment. In some cases, however, it can
be shown that the cerci really belong to an eleventh abdominal segment
which usually becomes fused with the tenth. With very few exceptions the
abdomen is without locomotor limbs. Paired processes on the eighth and
ninth abdominal segments may be specialized as external organs of
reproduction, but these are probably not appendages. The female genital
opening usually lies in front of the eighth abdominal segment, the male
duct opens on the ninth.
In all main points of their internal structure the Hexapoda agree with
other Arthropoda. Specially characteristic of the class, however, is the
presence of a complex system of air-tubes (tracheae) for respiration,
usually opening to the exterior by a series of paired spiracles on
certain of the body segments. The possession of a variable number of
excretory tubes (Malpighian tubes), which are developed as outgrowths of
the hind-gut and pour their excretion into the intestine, is also a
distinctive character of the Hexapoda.
The wings of insects are, in all cases, developed after hatching, the
younger stages being wingless, and often unlike the parent in other
respects. In such cases the development of wings and the attainment of
the adult form depend upon a more or less profound transformation or
metamorphosis.
With this brief summary of the essential characters of the Hexapoda, we
may pass to a more detailed account of their structure.
EXOSKELETON
The outer cellular layer (ectoderm or "hypodermis") of insects as of
other Arthropods, secretes a chitinous cuticle which has to be
periodically shed and renewed during the growth of the animal. The
regions of this cuticle have a markedly segmental arrangement, and the
definite hardened pieces (sclerites) of the exoskeleton are in close
contact with one another along linear sutures, or are united by
regions of the cuticle which are less chitinous and more membranous,
so as to permit freedom of movement.
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