_Excretory System._--Nitrogenous waste-matter is removed from the body
by the Malpighian tubes which open into the food-canal, usually where
the hind-gut joins the stomach. These tubes vary in number from four
to over a hundred in different orders of insects. The cells which line
them and also the cavities of the tubes contain urates, which are
excreted from the blood in the surrounding body-cavity. This cavity
contains an irregular mass of whitish tissue, the fat-body, consisting
of fat-cells which undergo degradation and become more or less filled
with urates. When the worn-out cells are broken down, the urates are
carried dissolved in the blood to the Malpighian tubes for excretion.
The fat-body is therefore the seat of important metabolic processes in
the hexapod body.
_Reproductive System._--All the Hexapoda are of separate sexes. The
ovaries (fig. 13) in the female are paired, each ovary consisting of a
variable number of tubes (one in the bristle-tail _Campodea_ and
fifteen hundred in a queen termite) in which the eggs are developed.
From each ovary an oviduct (fig. 13, od) leads, and in some of the
more primitive insects (bristle-tails, earwigs, may-flies) the two
oviducts open separately direct to the exterior. Usually they open
into a median vagina, formed by an ectodermal inpushing and lined with
chitin. The vagina usually opens in front of the eighth abdominal
sternite. Behind it is situated a spermatheca (fig. 14, sp) and the
ovipositor previously mentioned, with its three pairs of processes
(Fig. 14, G, g).
[Illustration: From Miall and Denny, _The Cockroach_, Lovell Reeve &
Co.
FIG. 13.--Ovaries of Cockroach, with Oviducts Od and Colleterial
Glands CG.]
[Illustration: From Miall and Denny, _The Cockroach_, Lovell Reeve &
Co.
FIG. 14.--Hinder Abdominal Segment and Ovipositor of Female Cockroach.
Magnified.
T^8 &c. Tergites.
S^7, 7th Sternite.
S^8, Sclerite between 7th and 8th sterna.
S^9, 8th Sclerite.
Od, Vagina.
sp, Spermatheca.
G, Anterior, and g, posterior gonapophyses.]
The paired testes of the male consist of a variable number of seminal
tubes, those of each testis opening into a _vas deferens_. In some
bristle-tails and may-flies, the two _vasa deferentia_ open
separately, but usually they lead into a sperm-reservoir, whence
issues a median ejaculatory duet. The male opening is on the ninth
abdominal segment, to which belong the processes that form the
claspers or genital armature. Accessory glands are commonly present in
connexion both with the male and the female reproductive organs. The
poison-glands of the sting in wasps and bees are well-known examples
of these.
EMBRYOLOGY
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