the bristle-tail _Lepisma_ and in dragon-flies they give rise to the
mid-gut. These views are not, however, supported by other recent
observers. J. Carriere's researches (1897) on the embryology of the
mason bee (_Chalicodoma_) agree entirely with the interpretations of
Kowalevsky and Heider, and so on the whole do those of F. Schwangart,
who has studied (1904) the embryonic development of Lepidoptera. He
finds that the endoderm arises from an anterior and a posterior
rudiment derived from the "endoblast," that many of the cells of these
rudiments wander into the yolk, and that the mesenteric epithelium
becomes reinforced by cells that migrate from the yolk. K. Escherich
(1901), after a new research on the embryology of the muscid Diptera,
claims that the fore and hind endodermal rudiments arise from the
blastoderm by invagination, and are from their origin distinct from
the mesoderm. On the whole it seems likely that the endoderm is
represented in part by the yolk, and in part by those anterior and
posterior rudiments which usually form the mesenteron, but that in
some Hexapoda the whole digestive tract may be ectodermal. It must be
admitted that some or the later work on insect embryology has
justified the growing scepticism in the universal applicability of the
"germ-layer theory." Heider has suggested, however, that the apparent
origin of the mid-gut from the stomodaeum and proctodaeum may be
explained by the presence of a "latent endoderm-group" in those
invaginations.
[Illustration: From Nussbaum in Miall and Denny, _The Cockroach_,
Lovell Reeve & Co.
FIG. 16.--Cross section of Embryo of German Cockroach
(_Phyllodromia_). S, serosa; A, amnion; E, ectoderm; N, rudiment of
nerve-cord; M, mesodermal pouches.]
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