_Forms of Larva._--The unlikeness of the young insect to its parent is
one of the factors that necessitates metamorphosis. It is instructive,
further, to trace among metabolic insects an increase in the degree of
this dissimilarity. An adult Hexapod is provided with a firm,
well-chitinized cuticle and six conspicuous jointed legs. Many larval
Hexapods might be defined in similar general terms, unlike as they are
to their parents in most points of detail. Examples of such are to be
seen in the grubs of may-flies, dragon-flies, lacewing-flies and
ground-beetles (fig. 24). This type of active, armoured larva--often
bearing conspicuous feelers on the head and long jointed cercopods on
the tenth abdominal segment--was styled campodeiform by F. Brauer
(1869), on account of its likeness in shape to the bristle-tail
_Campodea_. As an extreme contrast to this campodeiform type, we take
the maggot of the house-fly (fig. 25)--a vermiform larva, with soft,
white, feebly-chitinized cuticle and without either head-capsule or
legs. Between these two extremes, numerous intermediate forms can be
traced: the grub (wireworm) of a click-beetle, with narrow elongate
well-armoured body, but with the legs very short; the grub of a chafer,
with the legs fairly developed, but with the cuticle of all the
trunk-segments soft and feebly chitinized; the well-known caterpillar of
a moth (fig. 21, e) or saw-fly, with its long cylindrical body, bearing
the six shortened thoracic legs and a variable number of pairs of
"pro-legs" on the abdomen (this being the eruciform type of larva); the
soft, white, wood-boring grub of a longhorn-beetle or of the saw-fly
_Sirex_, with its stumpy vestiges of thoracic legs; the large-headed but
entirely legless, fleshy grub of a weevil; and the legless larva, with
greatly reduced head, of a bee. The various larvae of the above series,
however, have all a distinct head-capsule, which is altogether wanting
in the degraded fly maggot. These differences in larval form depend in
part on the surroundings among which the larva finds itself after
hatching; the active, armoured grub has to seek food for itself and to
fight its own battles, while the soft, defenceless maggot is provided
with abundant nourishment. But in general we find that elaboration of
imaginal structure is associated with degradation in the nature of the
larva, eruciform and vermiform larvae being characteristic of the
highest orders of the Hexapoda, so that unlikeness between parent and
offspring has increased with the evolution of the class.
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