5. _Sporogony._--This method of reproduction has been described by E.
Metchnikoff in _Cunina_ and allied genera. In individuals either of the
male or female sex, germ-cells which are quite undifferentiated and
neutral in character, become amoeboid, and wander into the endoderm.
They divide each into two sister-cells, one of which--the spore--becomes
enveloped by the other. The spore-cell multiplies by division, while the
enveloping cell is nutrient and protective. The spore cell gives rise to
a "spore-larva," which is set free in the coelenteron and grows into a
medusa. Whether sporogony occurs also in the polyp or not remains to be
proved.
6. _Sexual Reproduction and Embryology._--The ovum of Hydromedusae is
usually one of a large number of oögonia, and grows at the expense of
its sister-cells. No regular follicle is formed, but the oöcyte absorbs
nutriment from the remaining oögonia. In _Hydra_ the oöcyte is a large
amoeboid cell, which sends out pseudopodia amongst the oögonia and
absorbs nutriment from them. When the oöcyte is full grown, the residual
oögonia die off and disintegrate.
[Illustration: FIG. 47.--Budding from the Ectoderm (germinal epithelium)
in _Margellium_. (After C. Chun.)
A, The epithelium becomes two-layered.
B, The lower layer forms a solid mass of cells, which (C) becomes a
vesicle, the future endoderm, containing the coelenteric cavity
(coel), while the outer layer furnishes the future ectoderm.
D, E, F, a thickening of the ectoderm on the distal side of the bud
forms an entocodon (Gc).
G,H, Formation of the medusae.
s.c, Sub-umbral cavity.
r.c, Radial canal.
st, Stomach, which in H acquires a secondary communication with the
digestive cavity of the mother.
cc, Circular canal.
v, Velum.
t, Tentacle.]
The spermatogenesis and maturation and fertilization of the germ-cells
present nothing out of the common and need not be described here. These
processes have been studied in detail by A. Brauer [2] for _Hydra_.
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