_Polypodium hydriforme_ Ussow is a fresh-water form parasitic on the
eggs of the sterlet. A "stolon" of unknown origin produces thirty-two
buds, which become as many _Polypodia_; each has twenty-four tentacles
and divides by fission repeated twice into four individuals, each with
six tentacles. The daughter-individuals grow, form the full number of
twenty-four tentacles and divide again. The polyps are free and walk
on their tentacles. See Ussow [54].
_Tetraplatia volitans_ Viguier is a remarkable floating marine form.
See C. Viguier [56] and Delage and Hérouard (Hydrozoa [2]).
_Haleremita_ Schaudinn. See F. Schaudinn [50] and Delage and Hérouard
(HYDROZOA [2]).
In all the above-mentioned genera, with the exception of _Hydra_, the
life-cycle is so imperfectly known that their true position cannot be
determined in the present state of our knowledge. They may prove
eventually to belong to other orders. Hence only the genus _Hydra_ can
be considered as truly representing the order Eleutheroblastea. The
phylogenetic position of this genus has been discussed above.
ORDER II. Hydroidea seu Leptolinae.--Hydromedusae with alternation of
generations (metagenesis) in which a non-sexual polyp-generation
(trophosome) produces by budding a sexual medusa-generation (gonosome).
The polyp may be solitary, but more usually produces polyps by budding
and forms a polyp-colony. The polyp usually has the body distinctly
divisible into hydranth, hydrocaulus and hydrorhiza, and is usually
clothed in a perisarc. The medusae may be set free or may remain
attached to the polyp-colony and degenerate into a gonophore. When fully
developed the medusa is characterized by the sense organs being composed
entirely of ectoderm, developed independently of the tentacles, and
innervated from the sub-umbral nerve-ring.
The two kinds of persons present in the typical Hydroidea make the
classification of the group extremely difficult, for reasons explained
above. Hence the systematic arrangement that follows must be
considered purely provisional. A natural classification of the
Hydroidea has yet to be put forward. Many genera and families are
separated by purely artificial characters, mere shelf-and-bottle
groupings devised, for the convenience of the museum curator and the
collector. Thus many subdivisions are diagnosed by setting free
medusae in one case, or producing gonophores in another, although it
is very obvious, as pointed out above, that a genus producing medusae
may be far more closely allied to one producing gonophores than to
another producing medusae, or vice versa, and that in some cases the
production of medusae or gonophores varies with the season or the sex.
Moreover, P. Hallez [22] has recently shown that hydroids hitherto
regarded as distinct species are only forms of the same species grown
under different conditions.
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