SUB-ORDER 2. NARCOMEDUSAE.--Margin of the umbrella-lobed, tentacles
arising from the ex-umbrella at some distance from the margin;
tentaculocysts exposed, not enclosed in vesicles; gonads on the
sub-umbral floor of the stomach or of the gastric pouches.
[Illustration: FIG. 66.--_Cunina rhododactyla_, one of the
_Narcomedusae_. (After Haeckel.)
c, Circular canal.
h, "Otoporpae" or centripetal process of the marginal cartilaginous
ring connected with tentaculocyst.
k, Stomach.
l, Jelly of the disk.
r, Radiating canal (pouch of stomach).
tt, Tentacles.
tw, Tentacle root.]
The Narcomedusae exhibit peculiarities of form and structure which
distinguish them at once from all other Hydromedusae. The umbrella is
shallow and has the margin supported by a rim of thickened ectoderm, as
in the Trachomedusae, but not so strongly developed. The tentacles are
not inserted on the margin of the umbrella, but arise high up on the
ex-umbral surface, and the umbrella is prolonged into lobes
corresponding to the interspaces between the tentacles. The condition of
things can be imagined by supposing that in a medusa primitively of
normal build, with tentacles at the margin, the umbrella has grown down
past the insertion of the tentacles. As a result of this extension of
the umbrellar margin, all structures belonging to this region, namely,
the ring-canal, the nerve-rings, and the rim of thickened ectoderm, do
not run an even course, but are thrown into festoons, caught up under
the insertion of each tentacle in such a way that the ring-canal and its
accompaniments form in each notch of the umbrellar margin an inverted V,
the apex of which corresponds to the insertion of the tentacle; in some
cases the limbs of the V may run for some distance parallel to one
another, and may be fused into one, giving a figure better compared to
an inverted Y. Thus the ectodermal rim runs round the edge of each lobe
of the umbrella and then passes upwards towards the base of the tentacle
from the re-entering angle between two adjacent lobes, to form with its
fellow of the next lobe a tentacle-clasp or _peronium_, i.e. a streak of
thickened ectoderm supporting the tentacle. Similarly the ring-canal
runs round the edge of the lobe as the so-called festoon-canal, and then
runs upwards under the peronium to the base of the tentacle as one of a
pair of peronial canals, the limbs of the V-like figure already
mentioned. The nerve-rings have a similar course. The tentaculocysts are
implanted round the margins of the lobes of the umbrella and may be
supported by prolongations of the ectodermal rim termed _otoporpae_
(_Gehörspangen_). The radial canals are represented by wide gastric
pouches, and may be absent, so that the tentacles arise directly from
the stomach (_Solmaridae_). The tentacles are always solid, as in
Trachomedusae.
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