Both these medusae have sense-organs of a peculiar type, which are
said to contain an endodermal axis like the sense-organs of
Trachylinae, but the fact has recently been called in question for
_Limnocodium_ by S. Goto, who considers the genus to be allied to
_Olindias_. Allman, on the other hand, referred _Limnocodium_ to the
Leptomedusae.
In this connexion must be mentioned, finally, the medusae budded from
the fresh-water polyp _Microhydra_. The polyp-stages of _Limnocodium_
and _Microhydra_ are extremely similar in character. In both cases the
hydranth is extremely reduced and has no tentacles, and the polyp
forms a colony by budding from the base. In _Limnocodium_ the body
secretes a gelatinous mucus to which adhere particles of mud, &c.,
forming a protective covering. In _Microhydra_ no such protecting case
is formed. In view of the great resemblance between _Microhydra_ and
the polyp of _Limnocodium_, it might be expected that the medusae to
which they give origin would also be similar. As yet, however, the
medusa of _Microhydra_ has only been seen in an immature condition,
but it shows some well-marked differences from _Limnocodium_,
especially in the structure of the tentacles, which furnish useful
characters for distinguishing species amongst medusae. The possession
of a polyp-stage by _Limnocodium_ and _Microhydra_ furnishes an
argument against placing them in the Trachylinae. Their sense-organs
require renewed investigations. (Browne [10] and [10a].)
ORDER VI. Siphonophora.--Pelagic floating Hydrozoa with great
differentiation of parts, each performing a special function; generally
regarded as colonies showing differentiation of individuals in
correspondence with a physiological division of labour.
[Illustration: FIG. 68.--Diagram showing possible modifications of
medusiform and hydriform persons of a colony of _Siphonophora_. The
thick black line represents endoderm, the thinner line ectoderm. (After
Allman.)
n, Pneumatocyst.
k, Nectocalyces (swimming bells).
l, Hydrophyllium (covering-piece).
i, Generative medusiform person.
g, Palpon with attached palpacle, h.
e, Siphon with branched grappling tentacle, f.
m, Stem.]
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