[Illustration: FIG. 3.--Diagram showing the change of the Actinula (A)
into a Polyp (B); _a-b_, principal (vertical) axis; _c-d_, horizontal
axis. The endoderm is shaded, the ectoderm is left clear.]
It is convenient to distinguish two types of polyp by the names hydro
polyp and anthopolyp, characteristic of the Hydrozoa and Anthozoa
respectively. In the hydropolyp the body is typically elongated, the
height of the column being far greater than the diameter. The
peristome is relatively small and the mouth is generally raised on a
projecting spout or _hypostome_. The ectoderm loses entirely the
ciliation which it had in the planula and actinula stages and commonly
secretes on its external surface a protective or supporting
investment, the perisarc. Contrasting with this, the anthopolyp is
generally of squat form, the diameter often exceeding the height; the
peristome is wide, a hypostome is lacking, and the ectoderm, or so
much of it as is exposed, i.e. not covered by secretion of skeletal or
other investment, retains its ciliation throughout life. The internal
structural differences are even more characteristic. In the hydropolyp
the blastopore of the embryo forms the adult mouth situated at the
extremity of the hypostome, and the ectoderm and endoderm meet at this
point. In the anthopolyp the blastopore is carried inwards by an
in-pushing of the body-wall of the region of the peristome, so that
the adult mouth is an opening leading into a short ectodermal
oesophagus or stomodaeum, at the bottom of which is the blastopore.
Further, in the hydropolyp the digestive cavity either remains simple
and undivided and circular in transverse section, or may show ridges
projecting internally, which in this case are formed of endoderm
alone, without any participation of the mesogloea. In the anthopolyp,
on the other hand, the digestive cavity is always subdivided by
so-called mesenteries, in-growths of the endoderm containing vertical
lamellae of mesogloea (see ANTHOZOA). In short, the hydropolyp is
characterized by a more simple type of organization than the
anthopolyp, and is in most respects less modified from the actinula
type of structure.
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