The legs of Hymenoptera are of the typical insectan form, and the foot
is usually composed of five segments. In many families the trochanter
appears to be represented by two small segments, there being thus an
extra joint in the leg. It is almost certain that the distal of these
two segments really belongs to the thigh, but the ordinary
nomenclature will be used in the present article, as this character is
of great importance in discriminating families, and the two segments
in question are referred to the trochanter by most systematic writers.
[Illustration: After C. Janet, _Aiguillon de la Myrmica rubra_ (Paris,
1898).
FIG. 5.--Ovipositor or Sting of Red Ant (_Myrmica rubra_) Queen.
Magnified. The right sheath C (outer process of the ninth abdominal
segment--9) is shown in connexion with the guide B formed by the inner
processes of the 9th segment. The stylet A (process of the 8th
abdominal segment--8) is turned over to show its groove a, which works
along the tongue or rail b.]
The typical insectan ovipositor, so well developed among the
Hymenoptera, consists of three pairs of processes (gonapophyses) two
of which belong to the ninth abdominal segment and one to the eighth.
The latter are the cutting or piercing stylets (fig. 5, A) of the
ovipositor, while the two outer processes of the ninth segment are
modified into sheaths or feelers (fig. 5, C) and the two inner
processes form a guide (fig. 5, B) on which the stylets work, tongues
or rails on the "guide" fitting accurately into longitudinal grooves
on the stylet. In the different families of the Hymenoptera, there are
various modifications of the ovipositor, in accord with the habits of
the insects and the purposes to which the organ is put. The sting of
wasps, ants and bees is a modified ovipositor and is used for
egg-laying by the fertile females, as well as for defence. Most male
Hymenoptera have processes which form claspers or genital armature.
These processes are not altogether homologous with those of the
ovipositor, being formed by inner and outer lobes of a pair of
structures on the ninth abdominal segment.
Many points of interest are to be noted in the internal structure of
the Hymenoptera. The gullet leads into a moderate-sized crop, and
several pairs of salivary glands open into the mouth. The crop is
followed by a proventriculus which, in the higher Hymenoptera, forms
the so-called "honey stomach," by the contraction of whose wails the
solid and liquid food can be separated, passed on into the digestive
stomach, or held in the crop ready for regurgitation into the mouth.
Behind the digestive stomach are situated, as usual, intestine and
rectum, and the number of kidney (Malpighian) tubes varies from only
six to over a hundred, being usually great.
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