Most Hymenoptera are of moderate or small size, the giants of the
order--certain saw-flies and tropical digging-wasps--never reach the bulk
attained by the largest beetles, while the wing-spread is narrow compared
with that of many dragon-flies and moths. On the other hand, there are
thousands of very small species, and the tiny "fairy-flies"
(_Mymaridae_), whose larvae live as parasites in the eggs of various
insects, are excessively minute for creatures of such complex
organization. Hymenoptera are probably less widely distributed than
Aptera, Coleoptera or Diptera, but they are to be found in all except the
most inhospitable regions of the globe. The order is, with few
exceptions, terrestrial or aerial in habit. Comparatively only a few
species are, for part of their lives, denizens of fresh water; these, as
larvae, are parasitic on the eggs or larvae of other aquatic insects, the
little hymenopteron, _Polynema natans_, one of the "fairy-flies"--swims
through the water by strokes of her delicate wings in search of a
dragon-fly's egg in which to lay her own egg, while the rare _Agriotypus_
dives after the case of a caddis-worm. It is of interest that the waters
have been invaded by the parasitic group of the Hymenoptera, since in
number of species this is by far the largest of the order. No group of
terrestrial insects escapes their attacks--even larvae boring in wood are
detected by ichneumon flies with excessively long ovipositors. Not a few
cases are known in which a parasitic larva is itself pierced by the
ovipositor of a "hyperparasite," and even the offspring of the latter may
itself fall a victim to the attack of a "tertiary parasite."
_Fossil History._--Very little is known of the history of the
Hymenoptera previous to the Tertiary epoch, early in which, as we know
from the evidence of many Oligocene and Miocene fossils, all the more
important families had been differentiated. Fragments of wings from
the Lias and Oolitic beds have been referred to ants and bees, but the
true nature of these remains is doubtful.
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