In such a colony of connected individuals, the exact limits of the
separate "persons" are not always clearly marked out. Hence it is
necessary to distinguish between, first, the "zooids," indicated in
the case of the polyps by the hydranths, each with mouth and
tentacles; and, secondly, the "coenosarc," or common flesh, which
cannot be assigned more to one individual than another, but consists
of a more or less complicated network of tubes, corresponding to the
hydrocaulus and hydrorhiza of the primitive independent
polyp-individual. The coenosarc constitutes a system by which the
digestive cavity of any one polyp is put into communication with that
of any other individual either of the trophosome or gonosome. In this
manner the food absorbed by one individual contributes to the welfare
of the whole colony, and the coenosarc has the function of circulating
and distributing nutriment through the colony.
The hydroid colony shows many variations in form and architecture
which depend simply upon differences in the methods in which polyps
are budded.
[Illustration: After Hincks, Forbes, and Browne. A and B modified from
Hincks; C modified from Forbes's _Brit. Naked-eyed Medusae_.
FIG. 11.--_Lar sabellarum_ and two stages of its Medusa, _Willia
stellata_. A, colony of _Lar_; B and C, young and adult medusae.]
In the first place, buds may be produced only from the hydrorhiza,
which grows out and branches to form a basal _stolon_, typically
net-like, spreading over the substratum to which the founder-polyp
attached itself. From the stolon the daughter-polyps grow up
vertically. The result is a spreading or creeping colony, with the
coenosarc in the form of a root-like horizontal network (fig. 5, B;
11, A). Such a colony may undergo two principal modifications. The
meshes of the basal network may become very small or virtually
obliterated, so that the coenosarc becomes a crust of tubes tending to
fuse together, and covered over by a common perisarc. Encrusting
colonies of this kind are seen in _Clava squamata_ (fig. 5, A) and
_Hydractinia_ (figs. 9, 10), the latter having the perisarc calcified.
A further very important modification is seen when the tubes of the
basal perisarc do not remain spread out in one plane, but grow in all
planes forming a felt-work; the result is a massive colony, such as is
seen in the so-called Hydrocorallines (fig. 60), where the interspaces
between the coenosarcal tubes are filled up with calcareous matter, or
_coenosteum_, replacing the chitinous perisarc. The result is a stony,
solid mass, which contributes to the building up of coral reefs. In
massive colonies of this kind no sharp distinction can be drawn
between hydrorhiza and hydrocaulus in the coenosarc; it is practically
all hydrorhiza. Massive colonies may assume various forms and are
often branching or tree-like. A further peculiarity of this type of
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