As regards habit of life the vast majority of Hydromedusae are pelagic
organisms, floating on the surface of the open sea, propelling
themselves feebly by the pumping movements of the umbrella produced by
contraction of the sub-umbral musculature, and capturing their prey
with their tentacles. The genera _Cladonema_ (fig. 20) and
_Clavatella_ (fig. 21), however, are ambulatory, creeping forms,
living in rock-pools and walking, as it were, on the tips of the
proximal branches of each of the tentacles, while the remaining
branches serve for capture of food. _Cladonema_ still has the typical
medusan structure, and is able to swim about, but in _Clavatella_ the
umbrella is so much reduced, that swimming is no longer possible. The
remarkable medusa _Mnestra parasites_ is ecto-parasitic throughout
life on the pelagic mollusc _Phyllirrhoe_, attached to it by the
sub-umbral surface, and its tentacles have become rudimentary or
absent. It is interesting to note that _Mnestra_ has been shown by J.
W. Fewkes [15] and R. T. Günther [19] to belong to the same family
(_Cladonemidae_) as _Cladonema_ and _Clavatella_, and it is reasonable
to suppose that the non-parasitic ancestor of _Mnestra_ was, like the
other two genera, an ambulatory medusa which acquired louse-like
habits. In some species of the genus _Cunina_ (Narcomedusae) the
youngest individuals (actinulae) are parasitic on other medusae (see
below), but in later life the parasitic habit is abandoned. No other
instances are known of sessile habit in Hydromedusae.
[Illustration: After E. T. Browne, from _Proc. Zool. Soc. of London_.
FIG. 22.--_Corymorpha nutans_, adult female Medusa. Magnified 10
diameters.]
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